BA (1986) Natural Sciences, Zoology, Cambridge University Ph.D. (1990) MRC Molecular Neurobiology Unit, Cambridge University with Prof. Stephen Hunt 1990-1992: EMBO Long-term Fellowship, ZMBH, Uni Heidelberg with Prof. Peter H. Seeburg 1993-2000: Group Leader, MRC Laboratory of Molecular Biology, Cambridge 2001-2005: Group Leader, Clinical Neurobiology Dept, Uni Heidelberg August 2005: Professor of Neuroscience, University of Aberdeen