Abstract
Organic mulching is widely used in agriculture to improve soil conditions and enhance crop performance. However, mulch induces complex changes in soil processes, making process-based soil-crop models essential for understanding and predicting these effects. This non-systematic review summarizes the effects of mulch on soil hydrology, fertility, and temperature to evaluate how these processes are integrated into process-based soil-crop models. Current modeling approaches range from simple surface-residue representations that affect evaporation to more integrated frameworks that simulate decomposition and energy-balance feedbacks. Nevertheless, comprehensive and transferable representations of mulch effects remain limited. Future progress will require open-access datasets and modeling frameworks that explicitly represent mulch processes and feedback mechanisms with soil and crops. These improvements would enhance the implementation and optimization of mulch management strategies under varying environmental conditions.
1 What is organic mulching?
In natural ecosystems, plants influence the environment through the accumulation of litter, which stimulates soil life and strengthens ecosystem nutrient cycling (). In agriculture, most plant material is typically harvested, thus limiting the input of organic matter into the agroecosystem. Mimicking natural ecosystems in an agronomical context by applying undecomposed plant material, i.e., mulch, at least partially, compensates for crop residue removal. In this paper, the term ‘mulch’ refers specifically to organic mulching (using undecomposed plant-based litter) as opposed to plastic, inorganic, or living mulches.
Organic mulch is primarily used to improve the crop’s yield and water-use efficiency (WUE) (; ; ). A meta-analysis across 19 countries reported a 20% increase in yield and WUE for wheat and maize under straw mulching, though outcomes vary with environment, crop type, nitrogen and water availability (). Moreover, when matched to crop, management, and climate, mulch has been reported to buffer soil temperatures, support microbial activity and enhance nutrient cycling (; ; ). It has also been reported to reduce residual pesticides, fertilizers, heavy metals, while suppressing weeds (), and enhance disease tolerance (). However, the limited availability of local biomass or its conflict with other uses, and the labor-intensive nature and high transportation costs of its application are challenges to its broader adoption (; ).
A wide variety of organic mulch types have been used in agriculture, each with its own advantages in specific locations. The degree to which these mulches affect the soil quality, defined in this article as the soil’s capacity to function and support plant productivity (), depends on mulch type, thickness, and application method (e.g., full or partial cover and ridge–furrow systems) (; ; ). have provided a broad overview of application methods and mulch types. Readers are referred to this work for more in-depth information.
Although organic mulching is promoted as a valuable management strategy, its mechanisms and representation in soil-crop models remain insufficiently understood. Improving this representation is essential for evaluating mulch management strategies and supporting decision-making under varying environmental conditions. In this non-systematic review paper, we introduce the effects of organic mulching on soil hydrological processes, soil fertility, and soil temperature, focusing on mechanisms represented in process-based soil-crop models (Figure 1). This information is then used to evaluate how commonly process-based soil–crop models represent mulching, and to discuss their limitations and potential.
Figure 1
2 Effects of organic mulching on soil quality
2.1 Effects on soil hydrological processes
Runoff is the first pathway of water loss. Mulch protects the soil surface by dissipating raindrop kinetic energy, reducing particle detachment and flow velocity, and thereby limiting erosion and runoff (; ; ).
Mulches also increase soil water infiltration by limiting crust formation (; ; ) and increasing soil organic matter (SOM), which promotes soil aggregates and soil porosity (; ; ). This improved soil structure further enhances soil water retention capacity (; ; ; ). Although mulches intercept and absorb part of the rainfall (), field studies generally report improved soil water storage ().
Finally, by reducing solar radiation reaching the soil, mulching lowers soil evaporation and helps conserve moisture (; ; ; ). Over time, the application of mulch tends to improve the water budget for crops (; ).
2.2 Effects on soil fertility
Mulch influences soil fertility, defined as the soil’s ability to sustain plant growth by nutrient availability and biological activity (). Its effects on hydrological processes play a primary role, as nutrient transport is linked to transpiration, and drought can limit nutrient uptake despite the availability of nutrients (; ).
During its decomposition, mulch serves as a nutrient source, primarily for nitrogen (N). Overall, it improves crop N use efficiency () and reduces N and phosphate loss (; ; ; ). However, benefits depend on the synchronization of nutrient release with crop demand (). In this context, mulch quality in terms of C:N ratio is a key determinant of mineralization rate and the associated immobilization of soil nutrients (; ). Low C:N ratio mulches (e.g, leguminous residues) decompose faster, stimulate microbial mineralization, and enhance soil N availability, whereas high C:N ratio mulches (e.g, woody residues) immobilize N and reduce crop production (; ; ; ). The form of organic C also matters. Labile compounds, such as glucose cause stronger immobilization than more stable forms such as cellulose or tannins ().
By buffering moisture and providing carbon and nutrients, mulch stimulates microbial activity and accelerates residue turnover, enhancing nutrient cycling (; ; ; ) and improving SOM content (; ).
In summary, mulch can improve fertility by supplying nutrients and carbon and by maintaining soil moisture that supports microbial activity. The net outcome depends on mulch quality (e.g., C:N ratio and carbon form) and on how decomposition-driven nutrient release aligns with crop demand. Projected climate change will accelerate decomposition (), underscoring the importance of mulching on soil fertility.
2.3 Effects on soil temperature
Soil temperature is correlated with air temperature and surface cover () and governs the rate of various physical, chemical, and biological processes such as microbial activity, seed germination, plant development, pesticide degradation, and organic matter decomposition (; ). Mulch-driven changes in soil temperature have been observed to affect shoot growth, root development, biomass allocation, and nutrient uptake ().
Primarily, mulch alters surface energy fluxes by reflecting and absorbing shortwave radiation (; ; ; ), and by changing how energy is partitioned between latent and sensible heat during wetting and drying cycles (; ; ). In addition, a mulch layer acts as a thermal conductivity barrier, reducing the soil’s heat loss (; ; ). As a result, mulching buffers daily soil temperature fluctuations, cooling soils in summer and keeping them warm in winter, though outcomes depend on mulch thickness, mulch type, soil type, and climate (; ; ; ; ).
The effects of mulch-induced soil temperature changes on crop development depend on local climate, vary seasonally (), and are governed by crop-specific thermal requirements (; ). While a buffered soil temperature can improve and accelerate plant development under certain circumstances (; ), it can reduce yields or delay plant development under other circumstances (; ; ; ).
3 Representation of mulching in process-based soil-crop models
Mulching affects multiple interacting processes (Figure 1), making its impacts context-dependent. Process-based soil–crop models address this by representing soil–crop dynamics and interactions to simulate complex outcomes such as yield and soil water balance (). These models are widely used for decision support in crop management, studying genotype–environment interactions, and climate impact analysis. Consequently, process-based models can assess the effects of mulch when key processes are represented (). However, many models lack explicit mulch modules or have not been tested for this purpose.
3.1 General simulated soil processes
Models can be categorized into four distinct types according to the processes they represent. In this model, mulch is typically represented as an additional interacting component with specific properties.
Type 1 models only include the direct hydraulic impact of a mulch layer. Most commonly, this effect is implemented by reducing soil evaporation and runoff. For example, a mulch layer can reduce soil evaporation by acting as a sponge. It absorbs water and evaporates the intercepted water before the soil water is depleted (). Surface runoff, on the other hand, can be reduced by increasing simulated surface roughness, which promotes temporary ponding and enhances the infiltration rate (; ). These processes are often simplified using a “bucket” approach, in which mulch is represented as an intercepting reservoir with a fixed storage capacity (; ). Overall, this representation improves simulated plant water availability.
Type 2 models expand on Type 1 by incorporating mulch effects on the soil nutrient transformation and transport processes. Soil nutrient dynamics are generally represented by mass-balance equations. In these equations, nutrient pools change according to process-based rates influenced by soil temperature, soil moisture, and soil pH (; ). Therefore, mulch is represented via the mass balances and process rates governing the simulated system (; ). Nutrient availability influences root uptake, crop growth, and, indirectly, transpiration and soil water dynamics through feedbacks between biomass production and water demand ().
Type 3 models also build on Type 1 by incorporating the effect of mulch on the system’s energy balance rather than on the nutrient cycle. Soil temperature is commonly simulated using a heat-transport equation (; ), with mulch effects implemented through modified surface energy balance terms ().
Type 4 models combine all three major connections of Type 1, 2, and 3 models: (i) the effect on water relations, (ii) the nutrient cycle, and (iii) the energy balance. These models simultaneously simulate water, nutrient, and energy balances and their effects on crop growth, including interactions such as the influence of soil temperature on water and nutrient availability (). While this is a common feature in soil-crop models, and Type 4 can be used to model all connections between mulch and the soil-crop ecosystem (e.g., ), in practice, Type 4 is rarely tested due to its complex parametrization and the limited availability of datasets for model evaluation (; ; ; ; ; ; ; ; ; ; ) (Table 1).
Table 1
| Model | Water reations | Nutrient cycle | Energy balance |
|---|---|---|---|
| APSIM – type 4 | |||
| x | |||
| x | x | ||
| x | x | ||
| AquaCrop – type 1 | |||
| x | |||
| x | |||
| x | |||
| DSSAT – type 4 | |||
| x | x | ||
| x | x | ||
| x | |||
| HYDRUS 1D/SWAP–WOFOST – type 4 | |||
| x | x | ||
| x | x | ||
| x | |||
| STICS – type 4 | |||
| x | x | ||
| x | x | ||
| x | x | x | |
Five common soil–crop models, their type classification, and representative publications on mulch-effect representation.
The three categories water relations, nutrient cycling, and energy balance are marked with an “x” when the publication accounts for direct mulch-induced modifications. Model can in theory represent more than what they’ve been tested for.
In addition, soil biotic processes are not well represented in soil-crop models, even though they play an important role in soil quality (see section 2.2).
3.2 Common process-based soil-crop models
Numerous process-based soil-crop models have been developed over time, but their ability to simulate the effects of mulch varies. Here, we discuss six commonly used soil-crop models and their ability to represent mulching. These models are categorized according to the classification described above. APSIM, Aquacrop, DSSAT-CSM, HYDRUS 1-D, SWAP-WOFOST, and STICS. Table 1 summarizes the main characteristics and mulch-related functionalities. It is important to note that this list is not exhaustive, and other models can also be adapted to represent mulch as well, e.g., ARMOSA (), PASTIS (), RothC (), and nameless models ().
APSIM (Agricultural Production Systems Simulator) () can simulate systems for a diverse range of plant, animal, soil, climate, and management interactions, including organic mulch. The SurfaceOM module calculates the addition and decomposition of organic material at the soil surface, influencing soil evaporation and runoff. Carbon and nitrogen cycling are represented by complementary modules: SoilN simulates belowground transformations, while RESIDUE describes aboveground residue dynamics. Indirect effects of mulch, such as increased biomass production and associated changes in transpiration, are incorporated through SoilN processes (). However, APSIM can not reproduce temperature-induced delays in crop phenology, because it does not account for canopy temperature or mulch-driven soil temperature effects on crop growth (, ; ). To address this limitation, modified the temperature-time representation in the model.
AquaCrop simulates crop yield responses to water and is specifically developed where water is a major limiting factor for productivity (). The model is designed to be simple, robust, and accurate, requiring relatively few parameters and input variables. In AquaCrop, mulch only affects soil evaporation through two characteristics: soil cover fraction (0-100%) and mulch type. While this simplicity enables rapid assessment of mulch effects on soil–crop water relations, the model does not account for nutrient and energy balances or their interactions. Therefore, AquaCrop is useful when these processes are negligible or outside the research scope, while avoiding overparameterization (; ; ). However, while all models are simplifications of reality, in real-world applications, the nutrient and energy balance can rarely be neglected. As such, the use of AquaCrop to model mulch effects should be done with caution and careful consideration of the limitations of the model.
DSSAT (Decision Support System for Agrotechnology Transfer) simulates growth, development, and yield for a range of crops from coupled soil-plant-atmosphere processes (). Adding a mulch layer in DSSAT has a dual effect on soil evaporation: it stores and evaporates intercepted water, and buffers soil evaporation by meeting evaporative demand first. Furthermore, mulch representation promotes temporary ponding and increases infiltration, the latter represented by modified runoff coefficients and water storage limits (). Soil temperature is computed with a solar radiation and albedo-based module (), which can potentially capture mulch effects. With regard to nutrient dynamics, the CENTURY module () incorporates the effects of decomposing surface organic residue layers on soil organic matter and nutrient cycling. This decomposition is influenced by soil temperature and soil water content ().
Hydrus–1D () and SWAP () are finite element models capable of simulating water flow, as well as heat and solute transport. SWAP () can be coupled with WOFOST () to simulate complex crop growth (). Neither model includes a built-in option for a simulated mulch layer (). Mulch effects can nevertheless be represented by (i) adding an explicit surface layer with parametrized hydraulic and thermal properties (; ; ), or (ii) using simplified boundary-condition approaches (). While mulching effects on nutrient dynamics could, in theory, be captured by calibrating solute-transport parameters in the two aforementioned models, to the best of our knowledge, no studies to date have applied this adaptation.
STICS (Simulateur mulTIdisciplinaire pour les Cultures Standard) simulates the dynamic interactions among crops, soil, climate, and agricultural management modules () and provides one of the most integrated mulch implementations among the models discussed. A mulch layer directly affects soil evaporation by intercepting precipitation () and reducing surface runoff. Water intercepted by the layer evaporates (), adding moisture to the atmosphere, reducing crop transpiration, and impacting soil temperature. Soil evaporation is further reduced through the energy balance via the interception of solar radiation by the mulch layer (). Surface runoff is affected by mulch through the presence of soil surface obstacles (). As such, a mulch layer reduced runoff in proportion to the amount of mulch (t ha-1) that is applied to the soil, using a threshold of mulch presence above which no runoff occurred (; ). In addition, STICS computes mulch decomposition rates that depend on the type of plant residue. Decay of the mulch, in turn, affects the nutrient cycle by altering soil nitrogen availability ().
4 Challenges and opportunities in mulch modeling
Organic mulches improve soil quality and stabilize crop performance, but their effects are context-dependent and vary with soil type, plant species, mulch type, mulching method, and local climate. Therefore, understanding and predicting mulch effects under varying environmental conditions requires process-based soil–crop models that simulate coupled soil–plant processes and their feedback mechanisms (Figure 1).
While a wide range of models exists, explicit representation of mulch remains limited or absent.
In hard-coded models, mulch is often represented by modelers as an additional surface or adapted soil layer based on field and laboratory measurements (; ). The development of more flexible and modular open-source modeling frameworks facilitates the integration of a mulch module, as demonstrated by .
A major challenge remains the development of robust and transferable parameterizations of the mulch model. Mainly due to the extended range of mulch effects, contrasting environmental conditions, multiple management strategies, and different model representations. Progress in this area will require well-documented experimental datasets to support parameter estimation, calibration, sensitivity analysis, and independent model validation.
Model selection should also remain consistent and be guided by data availability and study objectives: simpler approaches, such as AquaCrop, may be adequate under limited-data conditions, whereas more complex models, such as STICS, require extensive calibration to reliably represent mulch-driven processes.
Future advances in mulch representation will improve the predictive capacity of soil–crop models and support the implementation and optimization of mulching as a sustainable agricultural management strategy. In particular, improved process representation could help quantify how long mulch-induced changes in nutrient availability persist, how different mulch types modify soil temperature, and how mulching alters crop resilience to drought in terms of timing, duration, and severity. Ultimately, such developments would support the optimization of mulch management strategies for specific crops and environmental conditions.
Statements
Author contributions
RW: Investigation, Writing – original draft, Writing – review & editing, Conceptualization. TD: Conceptualization, Investigation, Writing – review & editing, Writing – original draft. SG: Writing – review & editing. WC: Writing – review & editing. MD: Writing – original draft, Conceptualization, Investigation, Writing – review & editing. JS: Supervision, Investigation, Conceptualization, Writing – review & editing, Writing – original draft.
Funding
The author(s) declared that financial support was received for this work and/or its publication. This research is funded by the Agency for Agriculture and Fisheries of the Flemish government of Flanders, Belgium and is part of the Relance Plan ‘Vlaamse Veerkracht’ 2021.
Acknowledgments
We would like to thank the Agency for Agriculture and Fisheries of the Flemish government for funding the SoilStruct Project, under which this review was initiated.
Conflict of interest
The author(s) declared that this work was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest.
Generative AI statement
The author(s) declared that generative AI was used in the creation of this manuscript. Generative AI was used for language editing, sentence rephrasing, grammar checking and improving the clarity of the manuscript.
Any alternative text (alt text) provided alongside figures in this article has been generated by Frontiers with the support of artificial intelligence and reasonable efforts have been made to ensure accuracy, including review by the authors wherever possible. If you identify any issues, please contact us.
Publisher’s note
All claims expressed in this article are solely those of the authors and do not necessarily represent those of their affiliated organizations, or those of the publisher, the editors and the reviewers. Any product that may be evaluated in this article, or claim that may be made by its manufacturer, is not guaranteed or endorsed by the publisher.
References
1
AGROVOC (2021). AGROVOC (Rome: FAO). doi: 10.4060/cb2838en
2
AhirwarS.SubbaiahR.GuptaP.TiwariM. K.TrivediM. M.VaishnavP. (2023). Simulation of maize phenology and grain yield using DSSAT model. Int. J. Environ. Climate Change13, 2545–2556. doi: 10.9734/ijecc/2023/v13i92628
3
AmatoM. T.GiménezD. (2024). Predicting monthly near-surface soil temperature from air temperature and the leaf area index. Agric. For. Meteorol.345, 109838. doi: 10.1016/j.agrformet.2023.109838
4
AminM. G.MostofaS. M. M. M.HasanM. M.PervinW.SharminJ.HossainM. D. (2023). Plant–water relations in subtropical maize fields under mulching and organic fertilization. Agric. Water Manage.286, 108394. doi: 10.1016/j.agwat.2023.108394
5
BakrN.ElbanaT. A.ArceneauxA. E.ZhuY.WeindorfD. C.SelimH. M. (2015). Runoff and water quality from highway hillsides: Influence compost/mulch. Soil Tillage Res.150, 158–170. doi: 10.1016/j.still.2015.01.014
6
Balwinder-SinghGaydonD. S.HumphreysE.EberbachP. L. (2011). The effects of mulch and irrigation management on wheat in Punjab, India—evaluation of the APSIM model. Field Crops Res.124, 1–13. doi: 10.1016/j.fcr.2011.04.016
7
Balwinder-SinghHumphreysE.GaydonD. S.EberbachP. L. (2016). Evaluation of the effects of mulch on optimum sowing date and irrigation management of zero till wheat in central Punjab, India using APSIM. Field Crops Res.197, 83–96. doi: 10.1016/j.fcr.2016.08.016
8
BeaudoinN.LecharpentierP.Ripoche-WachterD.StrulluL.MaryB.LéonardJ.et al. (2023). STICS Soil-Crop Model: Conceptual Framework, Equations and Uses (Versailles: Éditions Quae).
9
BlanchyG.BragatoG.Di BeneC.JarvisN.LarsboM.MeurerK.et al. (2023). Soil and crop management practices and the water regulation functions of soils: A qualitative synthesis of meta-analyses relevant to European agriculture. SOIL9, 1–20. doi: 10.5194/soil-9-1-2023
10
BooteK. (2019). Advances in Crop Modelling for a Sustainable Agriculture (Cambridge: Burleigh Dodds Science Publishing).
11
BrissonN.GaryC.JustesE.RocheR.MaryB.RipocheD.et al. (2003). An overview of the crop model STICS. Eur. J. Agron.18, 309–332. doi: 10.1016/S1161-0301(02)00110-7
12
BristowK. L.HortonR. (1996). Modeling the impact of partial surface mulch on soil heat and water flow. Theor. Appl. Climatol.54, 85–96. doi: 10.1007/BF00863561
13
CaoY.HeZ.ZhuT.ZhaoF. (2021). Organic-C quality as a key driver of microbial nitrogen immobilization in soil: A meta-analysis. Geoderma383, 114784. doi: 10.1016/j.geoderma.2020.114784
14
CattanioJ. H.KuehneR.VlekP. L. G. (2008). Organic material decomposition and nutrient dynamics in a mulch system enriched with leguminous trees in the Amazon. Rev. Bras. Ciec. Do Solo32, 1073–1086. doi: 10.1590/S0100-06832008000300016
15
ChavesB.LéonardJ.FerchaudF.SchmatzR.RecousS.GiacominiS. J. (2024). Modelling decomposition of crop residue mulches and the associated N2O emissions in a no-till system in southern Brazil. Soil Tillage Res.240, 106065. doi: 10.1016/j.still.2024.106065
16
ChenS.ZhangX.PeiD.SunH.ChenS. (2007). Effects of straw mulching on soil temperature, evaporation and yield of winter wheat: Field experiments on the North China Plain. Ann. Appl. Biol.150, 261–268. doi: 10.1111/j.1744-7348.2007.00144.x
17
ChungS.HortonR. (1987). Soil heat and water flow with a partial surface mulch. Water Resour. Res.23, 2175–2186. doi: 10.1029/WR023i012p02175
18
CookH. F.ValdesG. S. B.LeeH. C. (2006). Mulch effects on rainfall interception, soil physical characteristics and temperature under Zea Mays L. Soil Tillage Res.91, 227–235. doi: 10.1016/j.still.2005.12.007
19
CramerM. D.HoffmannV.VerboomG. A. (2008). Nutrient availability moderates transpiration in Ehrharta Calycina. New Phytol.179, 1048–1057. doi: 10.1111/j.1469-8137.2008.02510.x
20
CrutchfieldD. A. (1984). “ The effect of wheat straw mulch on weed control and corn growth,” in ETD Collection for University of Nebraska-Lincoln, 1–200.
21
DöringT. F.BrandtM.HeßJ.FinckhM. R.SauckeH. (2005). Effects of straw mulch on soil nitrate dynamics, weeds, yield and soil erosion in organically grown potatoes. Field Crops Res.94, 238–249. doi: 10.1016/j.fcr.2005.01.006
22
DouY.ZhaoH.YangH.WangT.LiuG.WangZ.et al. (2024). The first factor affecting dryland winter wheat grain yield under various mulching measures: Spike number. J. Integr. Agric.23, 836–848. doi: 10.1016/j.jia.2023.05.034
23
DuC.LiL.XieJ.KhanK. S.WangL.LuoZ.et al. (2026). Endophytic fungi-mediated drought tolerance in spring wheat under no-tillage with straw cover on the Loess Plateau, China. Soil Tillage Res.256, 106857. doi: 10.1016/j.still.2025.106857
24
DusserreJ.AutfrayP.RakotoariveloM.RakotosonT.RaboinL.-M. (2020). Effects of contrasted cropping systems on yield and N balance of upland rainfed rice in Madagascar: Inputs from the DSSAT model. Exp. Agric.56, 355–370. doi: 10.1017/S0014479720000010
25
El-BeltagiH. S.BasitA.MohamedH. I.AliI.UllahS.KamelE. A.et al. (2022). Mulching as a sustainable water and soil saving practice in agriculture: A review. Agronomy12, 1881. doi: 10.3390/agronomy12081881
26
EnriqueG.BraudI.Jean-LouisT.MichelV.PierreB.Jean-ChristopheC. (1999). Modelling heat and water exchanges of fallow land covered with plant-residue mulch. Agric. For. Meteorol.97, 151–169. doi: 10.1016/S0168-1923(99)00081-7
27
Estrella DelgadoD. C.De SwaefT.VanderborghtJ.El MouttaqiA.HirichA.GarréS. (2026). Evaluating management strategies with crop modeling for salinity stress mitigation in quinoa: A case study from Laayoune, Morocco. Agric. Water Manage.325, 110207. doi: 10.1016/j.agwat.2026.110207
28
FanD.JiaG.WangY.YuX. (2023). The effectiveness of mulching practices on water erosion control: A global meta-analysis. Geoderma438, 116643. doi: 10.1016/j.geoderma.2023.116643
29
FAO (2021). Recarbonizing Global Soils: A Technical Manual of Recommended Sustainable Soil Management (Rome: FAO). doi: 10.4060/cb6595en
30
FindelingA.GarnierP.CoppensF.LafolieF.RecousS. (2007). Modelling water, carbon and nitrogen dynamics in soil covered with decomposing mulch. Eur. J. Soil Sci.58, 196–206. doi: 10.1111/j.1365-2389.2006.00826.x
31
FortinM.‐.PierceF. J. (1990). Developmental and growth effects of crop residues on corn. Agron. J.82, 710–715. doi: 10.2134/agronj1990.00021962008200040013x
32
FragaH.SantosJ. A. (2018). Vineyard mulching as a climate change adaptation measure: Future simulations for Alentejo, Portugal. Agric. Syst.164, 107–115. doi: 10.1016/j.agsy.2018.04.006
33
GavitoM. E.CurtisP. S.MikkelsenT. N.JakobsenI. (2001). Interactive effects of soil temperature, atmospheric carbon dioxide and soil N on root development, biomass and nutrient uptake of winter wheat during vegetative growth. J. Exp. Bot.52, 1913–1923. doi: 10.1093/jexbot/52.362.1913
34
GaydonD. S.Balwinder-SinghWangE.PoultonP. L.AhmadB.AhmedF.et al. (2017). Evaluation of the APSIM model in cropping systems of Asia. Field Crops Res.204, 52–75. doi: 10.1016/j.fcr.2016.12.015
35
GentileR.VanlauweB.SixJ. (2011). Litter quality impacts short- but not long-term soil carbon dynamics in soil aggregate fractions. Ecol. Appl.21, 695–703. doi: 10.1890/09-2325.1
36
GijsmanA. J.HoogenboomG.PartonW. J.KerridgeP. C. (2002). Modifying DSSAT crop models for low‐input agricultural systems using a soil organic matter–residue module from CENTURY. Agron. J.94, 462–474. doi: 10.2134/agronj2002.4620
37
GroenendijkP.BoogaardH.HeinenM.KroesJ. G.SupitI.De WitA. (2016). Simulation Nitrogen-Limited Crop Growth With Swap/Wofost : Process Descriptions and User Manual (Wageningen: Wageningen Environmental Research). doi: 10.18174/400458
38
GusevY. M.DzhoganL. Y.NasonovaO. N. (2018). Modelling the impact of mulching the soil with plant remains on water regime formation, crop yield and energy costs in agricultural ecosystems. Proc. Int. Assoc. Hydrol. Sci.376, 77–82. doi: 10.5194/piahs-376-77-2018
39
HeinenM.MulderM.Van DamJ.BartholomeusR.De Jong Van LierQ.De WitJ.et al. (2024). SWAP 50 years: Advances in modelling soil-water-atmosphere-plant interactions. Agric. Water Manage.298, 108883. doi: 10.1016/j.agwat.2024.108883
40
HillelD. I. (1998). Environmental Soil Physics (Waltham: Academic press).
41
HoogenboomG.PorterC. H.BooteK. J.SheliaV.WilkensP. W.SinghU.et al. (2019). “ The DSSAT crop modeling ecosystem,” in Advances in Crop Modelling for a Sustainable Agriculture (Cambridge: Burleigh Dodds Science Publishing). doi: 10.19103/AS.2019.0061.10
42
HuN.WangX.PangL.LuJ.YangJ.XiaoX.et al. (2026). Ecological trade-offs of plastic film and straw mulching: Mechanistic insights from soil structure and carbon–nitrogen. Agronomy16, 470. doi: 10.3390/agronomy16040470
43
IqbalR.RazaM. A. S.ValipourM.SaleemM. F.ZaheerM. S.AhmadS.et al. (2020). Potential agricultural and environmental benefits of mulches—a review. Bull. Natl. Res. Cent.44, 75. doi: 10.1186/s42269-020-00290-3
44
IrkisoA. B.KuhwaldM.ThiekenA. H.GreveP.ChemuraA. (2025). Application of DSSAT model to evaluate the effects of tillage methods on soil water balance during drought period. Model. Earth Syst. Environ.11, 200. doi: 10.1007/s40808-025-02334-x
45
JonesJ. W.HoogenboomG.PorterC. H.BooteK. J.BatchelorW. D.HuntL. A.et al. (2003). The DSSAT cropping system model. Eur. J. Agron.18, 235–265. doi: 10.1016/S1161-0301(02)00107-7
46
JonesJ. W.TsujiG. Y.HoogenboomG.HuntL. A.ThorntonP. K.WilkensP. W.et al. (1998). “ Decision support system for agrotechnology transfer: DSSAT V3,” in Understanding Options for Agricultural Production, vol. 7 . Eds. TsujiG. Y.HoogenboomG.ThorntonP. K. (Dordrecht: Springer Netherlands). doi: 10.1007/978-94-017-3624-4_8
47
JordánA.ZavalaL. M.GilJ. (2010). Effects of mulching on soil physical properties and runoff under semi-arid conditions in southern Spain. CATENA81, 77–85. doi: 10.1016/j.catena.2010.01.007
48
JuryW. A.GardnerW. R.GardnerW. H. (1991). Soil Physics. 5th ed (New York: J. Wiley).
49
KaderM. A.NakamuraK.SengeM.MojidM. A.KawashimaS. (2019). Numerical simulation of water- and heat-flow regimes of mulched soil in rain-fed soybean field in central Japan. Soil Tillage Res.191, 142–155. doi: 10.1016/j.still.2019.04.006
50
KaderM. A.SengeM.MojidM. A.ItoK. (2017). Recent advances in mulching materials and methods for modifying soil environment. Soil Tillage Res.168, 155–166. doi: 10.1016/j.still.2017.01.001
51
KarlenD. L.MausbachM. J.DoranJ. W.ClineR. G.HarrisR. F.SchumanG. E. (1997). Soil quality: a concept, definition, and framework for evaluation (a guest editorial). Soil Sci. Soc Am. J.61, 4–10. doi: 10.2136/sssaj1997.03615995006100010001x
52
KisakaM. O.Mucheru-MunaM.NgetichF. K.MugweJ. N.MugendiD. N.MairuraF.et al. (2016). Using APSIM-model as a decision-support-tool for long-term integrated-nitrogen-management and maize productivity under semi-arid conditions in Kenya. Exp. Agric.52, 279–299. doi: 10.1017/S0014479715000095
53
KisingaB.TherhaagE.KlaukB.GrossJ.Schulte-GeldermannE.DöringT. F. (2026). Mulch-driven tolerance in potatoes: a climate adaptation tool for enhanced yields and plant defence under drought stress conditions. Potato Res.69, 67. doi: 10.1007/s11540-026-10012-3
54
KodešováR.FérM.KlementA.NikodemA.TepláD.NeubergerP.et al. (2014). Impact of various surface covers on water and thermal regime of technosol. J. Hydrol.519, 2272–2288. doi: 10.1016/j.jhydrol.2014.10.035
55
KomariahItoK.SengeM.AdomakoJ. T.Afandi (2008). The influences of organic mulches on soil moisture content and temperatures: a case study of tapioca wastes application. J. Rainwater Catchment Syst.14, 1–8. doi: 10.7132/jrcsa.kj00004978336
56
KosternaE. (2014). The effect of covering and mulching on the soil temperature, growth and yield of tomato. Folia Hortic.26, 91–101. doi: 10.2478/fhort-2014-0009
57
KroesJ. G.Van DamJ. C.BartholomeusR. P.GroenendijkP.HeinenM.HendriksR. F. A.et al. (2017). SWAP version 4 environmental research. (Wageningen: Wageningen Environmental Research). doi: 10.18174/416321
58
LarssonL.BåthA. (1996). Evaluation of soil temperature moderating and moisture conserving effects of various mulches during a growing season. Acta Agriculturae Scandinavica Section B. - Soil Plant Sci.46, 153–160. doi: 10.1080/09064719609413128
59
LiQ.LiH.ZhangL.ZhangS.ChenY. (2018). Mulching improves yield and water-use efficiency of potato cropping in Chine: a meta-analysis. Field Crops Res.221, 50–60. doi: 10.1016/j.fcr.2018.02.017
60
LiuR.ThomasB. W.ShiX.ZhangX.WangZ.ZhangY. (2021). Effects of ground cover management on improving water and soil conservation in tree crop systems: a meta-analysis. CATENA199, 105085. doi: 10.1016/j.catena.2020.105085
61
LopesI.De LimaJ. L. M. P.MontenegroA. A. A.De CarvalhoA. A. (2025). Assessment of water retention and absorption of organic mulch under simulated rainfall for soil and water conservation. Soil Syst.9, 4. doi: 10.3390/soilsystems9010004
62
MahawarL.MishraA.TsitouriA.AlbrectsenB. R. (2026). Straw mulching differentially shapes the structure and function of below‐ground bacterial communities in potato depending on eDNA source and cultivar. Plant-Environ. Interact.7, e70131. doi: 10.1002/pei3.70131
63
McCownR. L.HammerG. L.HargreavesJ. N. G.HolzworthD. P.FreebairnD. M. (1996). APSIM: a novel software system for model development, model testing and simulation in agricultural systems research. Agric. Syst.50, 255–271. doi: 10.1016/0308-521X(94)00055-V
64
MoriA. S.CornelissenJ. H. C.FujiiS.OkadaK.IsbellF. (2020). A meta-analysis on decomposition quantifies afterlife effects of plant diversity as a global change driver. Nat. Commun.11, 4547. doi: 10.1038/s41467-020-18296-w
65
MulumbaL. N.LalR. (2008). Mulching effects on selected soil physical properties. Soil Tillage Res.98, 106–111. doi: 10.1016/j.still.2007.10.011
66
MupangwaW.DimesJ.WalkerS.TwomlowS. (2011). Measuring and simulating maize (Zea mays L.) yield responses to reduced tillage and mulching under semi-arid conditions. Agric. Sci.02, 167–174. doi: 10.4236/as.2011.23023
67
Nazemi RafiZ.KazemiF. (2021). Effects of planting combinations and mulch types on soil moisture and temperature of xeric landscapes. Urban Forestry Urban Greening58, 126966. doi: 10.1016/j.ufug.2020.126966
68
NovakM. D.ChenW.HaresM. A. (2000). Simulating the radiation distribution within a barley-straw mulch. Agric. For. Meteorol.102, 173–186. doi: 10.1016/S0168-1923(00)00096-4
69
PalmC.Blanco-CanquiH.DeClerckF.GatereL.GraceP. (2014). Conservation agriculture and ecosystem services: an overview. Agric. Ecosyst. Environ.187, 87–105. doi: 10.1016/j.agee.2013.10.010
70
PesceS.BaluganiE.De PazJ. M.MarazzaD.ViscontiF. (2024). A modified version of RothC to model the direct and indirect effects of rice straw mulching on soil carbon dynamics, calibrated in two Valencian citrus orchards. Soil Syst.8, 12. doi: 10.3390/soilsystems8010012
71
PorterC. H.JonesJ. W.AdikuS.GijsmanA. J.GargiuloO.NaabJ. B. (2010). Modeling organic carbon and carbon-mediated soil processes in DSSAT v4.5. Operational Res.10, 247–278. doi: 10.1007/s12351-009-0059-1
72
PriceJ.RochefortL.QuintyF. (1998). Energy and moisture considerations on cutover peatlands: surface microtopography, mulch cover and sphagnum regeneration. Ecol. Eng.10, 293–312. doi: 10.1016/S0925-8574(98)00046-9
73
ProbertM. E.DelveR. J.KimaniS. K.DimesJ. P. (2005). Modelling nitrogen mineralization from manures: representing quality aspects by varying C:N ratio of sub-pools. Soil Biol. Biochem.37, 279–287. doi: 10.1016/j.soilbio.2004.07.040
74
QinW.HuC.OenemaO. (2015). Soil mulching significantly enhances yields and water and nitrogen use efficiencies of maize and wheat: a meta-analysis. Sci. Rep.5, 16210. doi: 10.1038/srep16210
75
RanaivosonL.NaudinK.RipocheA.AffholderF.RabeharisoaL.CorbeelsM. (2017). Agro-ecological functions of crop residues under conservation agriculture. a review. Agron. Sustain. Dev.37, 26. doi: 10.1007/s13593-017-0432-z
76
RooseT.SchnepfA. (2008). Mathematical models of plant–soil interaction. Philos. Trans. R. Soc. A. Mathematical Phys. Eng. Sci.366, 4597–4611. doi: 10.1098/rsta.2008.0198
77
RossiG.BeniC.NeriU. (2024). Organic mulching: a sustainable technique to improve soil quality. Sustainability16, 10261. doi: 10.3390/su162310261
78
ScopelE.Da SilvaF. A. M.CorbeelsM.AffholderF.MarauxF. (2004). Modelling crop residue mulching effects on water use and production of maize under semi-arid and humid tropical conditions. Agronomy24, 383–395. doi: 10.1051/agro:2004029
79
SekhonN. K.HiraG. S.SidhuA. S.ThindS. S. (2005). Response of soyabean (Glycine max Mer.) to wheat straw mulching in different cropping seasons. Soil Use Manage.21, 422–426. doi: 10.1079/SUM2005356
80
ShanonoN. J.AbbaB. S.NasiDiN. M. (2022). Evaluation of Aqua-Crop model using onion crop under deficit irrigation and mulch in semi-arid Nigeria. Turkish J. Agric. Eng. Res.3, 131–145. doi: 10.46592/turkager.1078082
81
SharmaR.BhardwajS. (2017). Effect of mulching on soil and water conservation -a review. Agric. Rev.38. doi: 10.18805/ag.R-1732
82
ShirishaK.RayL. I. P.ParidaP. K.SinghA. K.SinghN. J. (2023). Organic mulching effects on water productivity, soil temperature, growth and yield of lentil varieties. Int. J. Bio-Resource Stress Manage.14, 1099–1107. doi: 10.23910/1.2023.3556c
83
ŠimůnekJ.Van GenuchtenM. T.ŠejnaM. (2008). Development and applications of the HYDRUS and STANMOD software packages and related codes. Vadose Zone J.7, 587–600. doi: 10.2136/vzj2007.0077
84
SokolN. W.SlessarevE.MarschmannG. L.NicolasA.BlazewiczS. J.BrodieE. L.et al. (2022). Life and death in the soil microbiome: how ecological processes influence biogeochemistry. Nat. Rev. Microbiol.20, 415–430. doi: 10.1038/s41579-022-00695-z
85
SouzaR.JhaA.CalabreseS. (2022). Quantifying the hydrological impact of soil mulching across rainfall regimes and mulching layer thickness. J. Hydrol.607, 127523. doi: 10.1016/j.jhydrol.2022.127523
86
StedutoP.HsiaoT. C.RaesD.FereresE. (2009). AquaCrop—the FAO crop model to simulate yield response to water: I. concepts and underlying principles. Agron. J.101, 426–437. doi: 10.2134/agronj2008.0139s
87
SuiH.ZengD.ChenF. (1992). A numerical model for simulating the temperature and moisture regimes of soil under various mulches. Agric. For. Meteorol.61, 281–299. doi: 10.1016/0168-1923(92)90054-8
88
TadielloT.GabbrielliM.BottaM.AcutisM.BechiniL.RagagliniG.et al. (2023). A new module to simulate surface crop residue decomposition: description and sensitivity analysis. Ecol. Modell.480, 110327. doi: 10.1016/j.ecolmodel.2023.110327
89
TruongT. H. H.MarschnerP. (2020). Plant residues differing in C/N ratio in mulch and soil — the effect of the mulch on nutrient availability and microbial biomass is more pronounced with higher leaching amount. Soil Ecol. Lett.2, 317–326. doi: 10.1007/s42832-020-0036-4
90
TuC.RistainoJ. B.HuS. (2006). Soil microbial biomass and activity in organic tomato farming systems: effects of organic inputs and straw mulching. Soil Biol. Biochem.38, 247–255. doi: 10.1016/j.soilbio.2005.05.002
91
UllahI.ShahS. T.BasitA.SajidM.ArifM.AhmadN.et al. (2022). “ Mulching: a new concept for climate smart agriculture,” in Mulching in Agroecosystems. Eds. AkhtarK.ArifM.RiazM.WangH. (Singapore: Springer Nature Singapore). doi: 10.1007/978-981-19-6410-7_17
92
UmeshB.ReddyK. S.PolisgowdarB. S.MaruthiV.SatishkumarU.AyyanagoudarM. S.et al. (2022). Assessment of climate change impact on (Zea mays L.) through AquaCrop model in semi-arid alfisol of southern Telangana. Agric. Water Manage.274, 107950. doi: 10.1016/j.agwat.2022.107950
93
VeenG. F. (.FryE. L.Ten HoovenF. C.KardolP.MorriënE.De LongJ. R. (2019). The role of plant litter in driving plant-soil feedbacks. Front. Environ. Sci.7, 168. doi: 10.3389/fenvs.2019.00168
94
WangX.FanJ.XingY.XuG.WangH.DengJ.et al. (2019). The effects of mulch and nitrogen fertilizer on the soil environment of crop plants. Adv. Agron.153. doi: 10.1016/bs.agron.2018.08.003
95
WoodwardJ. (1699). Some thoughts and experiments concerning vegetation. Philos. Trans.21, 193–227. doi: 10.1098/rstl.1699.0040
96
YanJ.WangL.HuY.TsangY. F.ZhangY.WuJ.et al. (2018). Plant litter composition selects different soil microbial structures and in turn drives different litter decomposition pattern and soil carbon sequestration capability. Geoderma319, 194–203. doi: 10.1016/j.geoderma.2018.01.009
97
YangY.-M.LiuX.-J.LiW.-Q.LiC.-Z. (2006). Effect of different mulch materials on winter wheat production in desalinized soil in Heilonggang region of North China. J. Zhejiang Univ. Sci. B.7, 858–867. doi: 10.1631/jzus.2006.b0858
98
YuY.-Y.TurnerN. C.GongY.-H.LiF.-M.FangC.GeL.-J.et al. (2018). Benefits and limitations to straw- and plastic-film mulch on maize yield and water use efficiency: a meta-analysis across hydrothermal gradients. Eur. J. Agron.99, 138–147. doi: 10.1016/j.eja.2018.07.005
99
ZhangW.-P.FornaraD.YangH.YuR.-P.CallawayR. M.LiL. (2023). Plant litter strengthens positive biodiversity-ecosystem functioning relationships over time. Trends Ecol. Evol.38, 473–484. doi: 10.1016/j.tree.2022.12.008
100
ZhangS.LovdahlL.GripH.TongY.YangX.WangQ. (2009). Effects of mulching and catch cropping on soil temperature, soil moisture and wheat yield on the loess plateau of China. Soil Tillage Res.102, 78–86. doi: 10.1016/j.still.2008.07.019
101
ZizingaA.MajaliwaJ. G.TietjenB.BedadiB.De SalesR. A.BeesigamukamaD. (2022). Simulating maize productivity under selected climate smart agriculture practices using AquaCrop model in a sub-humid environment. Sustainability14, 2036. doi: 10.3390/su14042036
Summary
Keywords
crop production, organic mulching, process-based soil-crop model, soil fertility, soil hydrological processes, soil temperature
Citation
Willemet R, De Swaef T, Garré S, Cornelis WM, De Boever M and Schreel JDM (2026) Mulch representation in soil-crop models. Front. Agron. 8:1900760. doi: 10.3389/fagro.2026.1900760
Received
05 June 2026
Revised
04 July 2026
Accepted
07 July 2026
Published
23 July 2026
Volume
8 - 2026
Edited by
Indra Jeet Chaudhary, Savitribai Phule Pune University, India
Reviewed by
Khuram Shehzad Khan, China Agricultural University, China
Pengliang Tian, Gansu Agricultural University, China
Updates
Copyright
© 2026 Willemet, De Swaef, Garré, Cornelis, De Boever and Schreel.
This is an open-access article distributed under the terms of the Creative Commons Attribution License (CC BY). The use, distribution or reproduction in other forums is permitted, provided the original author(s) and the copyright owner(s) are credited and that the original publication in this journal is cited, in accordance with accepted academic practice. No use, distribution or reproduction is permitted which does not comply with these terms.
*Correspondence: Rémy Willemet, Remy.Willemet@ilvo.vlaanderen.be
Disclaimer
All claims expressed in this article are solely those of the authors and do not necessarily represent those of their affiliated organizations, or those of the publisher, the editors and the reviewers. Any product that may be evaluated in this article or claim that may be made by its manufacturer is not guaranteed or endorsed by the publisher.