Abstract
This work presents evidence of double diffusive convection (DDC) in the form of salt fingers (SF) forming a thermohaline staircase in the Cretan Sea (South Aegean Sea). The phenomenon was identified at the eastern edge of the Cretan Sea in March 2023 but had vanished by September 2023. To estimate vertical diffusivities and fluxes, four DDC models were applied to the hydrographic data. The vulnerability of the staircase appears to be related to weak, downward, salt finger–induced density fluxes, possibly caused by tilting of the salt fingers due to prevailing strong horizontal shears. Two of the four model implementations showed closer agreement with predictions from a previous staircase study in the area. Analysis of the density ratio and Turner angle from individual cruises demonstrated the widespread susceptibility of the Cretan Sea water column to such phenomena. Specifically, cruise data from 2016 to 2023, covering the layer between the intermediate salinity maximum (Levantine Intermediate Water) and the deep salinity minimum (Transitional Mediterranean Water), showed that 71%–85% of Turner angle values lie in the salt fingering zone. Thermohaline staircases in the Cretan Sea play a significant role in shaping the physical and biochemical processes of the basin by transporting salt and density toward deeper layers while uplifting nutrients. Although small scale, the phenomenon can significantly influence the evolution of the water column and large-scale climatic processes, such as vertical heat and salt transport and overturning circulation in deep basins.
1 Introduction
1.1 Oceanographic setting
The Cretan Sea lies at the southern extremity of the Aegean Sea and constitutes the largest basin of the South Aegean Sea. It is bounded to the north by the Cyclades island complex and to the south by the island of Crete (Figure 1). This sea is a key Eastern Mediterranean (EMed) basin, acting both as a large heat and salt reservoir for the entire EMed and as a source of dense waters for the Levantine and Ionian basins (; ; ).
Figure 1
Its intermediate layers are occupied by a mixture of warm, saline waters either imported from the Levantine Sea through the eastern Cretan Straits [Levantine Intermediate Water (LIW)] or produced locally [Cretan Intermediate Water (CIW)]. Below these layers lies a colder, less saline water mass imported from the open EMed. The core of this mass, known as Transitional Mediterranean Water (TMW), is characterized by a local salinity and oxygen minimum. It enters the basin from both the eastern and western Cretan Straits at sill depths (~800–1,000 m) and stabilizes according to its typical potential density (σθ ~29.17–29.18 kg/m³) at horizons below the LIW/CIW layer. The TMW core depth varies interannually and conforms to the changing density stratification of the Cretan Sea below the intermediate waters (for information on Cretan Sea hydrology, see
Thus, the Cretan Sea is a basin where waters of different origins and characteristics converge and gradually mix. The aforementioned TMW enters through the Cretan Straits and forms a layer that separates the saltier CIW from the Cretan Deep Water (CDW). Studies have shown that the TMW signal (in the form of a salinity minimum) is strongest during maximal exchange through the Cretan Straits, and that when this exchange weakens, the TMW layer is gradually eroded through mixing with CIW and CDW (
1.2 Double diffusive convection and salt finger thermohaline staircases
In this work, we present evidence of a salt finger thermohaline staircase (SF-TS) in the eastern Cretan Sea. These staircases comprise a spatially organized evolution of double diffusive convection (DDC). DDC arises from the contact of two fluid mixtures whose density-defining thermodynamic properties undergo molecular diffusion at different rates (see
For seawater, double diffusive phenomena emerge because the molecular diffusivity ratio of heat () and salt () is on the order of 100 (). In the ocean, DDC rises in the stably stratified interior () of an oceanic basin, and emerges as a gradually enhancing instability within a water mass characterized by gradients of the same sign. Positive or negative gradients give rise to the two known DDC variants; salt fingers (SF, for positive gradients) and diffusive layers (DL, for negative gradients).
Generally, diffusive instability occurs when the unequal molecular fluxes of heat and salt create statically unstable vertical density profiles (inversions), leading to localized convective motions. For both DDC variants, the instability develops due to differences in the magnitude of molecular fluxes of heat and salt, which are controlled by the interaction between molecular diffusivities and the respective ambient gradients. . Under certain conditions, the instabilities evolve into spatially organized staircases, such as the SF_ThS herein presented. Salt fingering occurs in the presence of positive gradients , where both and increase upwards in the water column and has been first described and theoretically explained by M.E. Stern (
The growth of salt fingers halts when their height reaches a limiting value (e.g., ~30 cm, according to
The overall effective diffusive fluxes, from the rise of the initial growth of SF to their turbulent break down, can be described by the SF diffusivities of salt and potential temperature, . These diffusivities are unequal, with Indeed, the combination of the formulas for fluxes and , buoyancy flux ratio , density ratio in terms of the heat expansion and saline contraction coefficients, , and the background vertical gradients of salinity and potential temperature, , , yield a ratio of diffusivities, . This in turn, if combined with the flux ratio formula of Kunze’s model for growing fingers (
The salt-finger staircase studied herein comprises a spatiotemporally persistent and coherent configuration of the latter sequel, wherein ‘interfaces’, i.e. vertical regimes (1 to 10 m) of growing fingers with marked gradients are alternated by ‘layers’ (10 to 100m or more), that have been homogenized by the finger-into-turbulence breakouts (). Diffusive-layer staircases are quite analogous (
According to You (2002), the criteria for salt fingering are met in 30% of the world’s oceans, wherever warm, saline water masses overlie colder, less saline ones. In the Mediterranean Sea, favorable conditions for the onset of salt fingering exist throughout most of the basin, as warm and salty intermediate water masses of the upper thermohaline cell overlie the colder, less saline deep layers (
In contrast, observations of ThSs in the EMed are relatively scarce, reported mainly in the southeastern Ionian Sea in the vicinity of water lenses with and characteristics different from the surrounding masses (Zodiatis, 1992;
Finally, of particular importance to the present analysis is the value used to indicate the strength of SF_DDCs, namely the so-called density ratio, , where is the thermal expansion coefficient and the haline contraction coefficient. expresses the ratio of temperature to salinity contributions on density stratification and can take values from -∞ to +∞. It comprises a measure of the compensation between the temperature and salinity gradients on the density stratification. Strong fingering is more active at , while values correspond to weaker fingering. The closer is to 1, the stronger the fingers.
Apart from , the Turner angle (
The present work is divided into two main aims. The first is to estimate DDC fluxes and diffusivities produced by the recorded SF-TS and to compare them with previous results for the Cretan Sea, the EMed, and other regions. The second is to evaluate the susceptibility of the Cretan Sea to SF-DDC events by statistically analyzing the stability ratio and the Turner angle, based on an ensemble of spatiotemporal coverages within the basin.
2 Materials and methods
In March 2023, during a cruise with R/V AEGAEO of the Hellenic Centre for Marine Research (HCMR) in the Aegean Sea under the Marine Strategy Framework Directive (MSFD) monitoring program, a profile in the eastern Cretan Sea (red dot in Figure 1) was sampled with a shipborne conductivity–temperature–depth (CTD) probe. A well-defined SF-TS (Figure 2) was detected in this cast but not in the subsequent MSFD cast at the same location in September 2023, nor in a later cast obtained during the CLIMPACT project in April 2025. Apart from these CTD casts, additional data from the region were considered, obtained from: a) the CRELEV 2016 oceanographic cruise, conducted in June 2016 under EUROFLEETS2 with R/V AEGAEO (
Figure 2

CTD cast of March 17th, 2023. Cast position: latitude 35° 45.20 N, longitude 026° 13.71 E Potential temperature (theta) and salinity of the thermohaline staircase profile (a). Same parameters in the 650–1,000 dbar layer (b).
Glider data presented herein are part of HCMR’s glider mission no. 33, conducted north of Crete in the framework of repeated seasonal missions established in the Cretan Sea. The mission lasted 18 days (March 15–April 2, 2023), covering ~400 km mostly north of Crete and performing vertical measurements in a sawtooth pattern. An Alseamar SeaExplorer glider equipped with a Seabird GPCTD payload was used. Full data with a 4 s sampling period were retrieved directly upon recovery of the glider and subsequently interpolated to 1 dbar bins. These data were validated by comparison with reference CTD casts performed at both the beginning and the end of the mission. To construct Turner angle distributions from the dataset, a 10 dbar running mean filter was applied.
The criteria for delimiting the layers followed those of
Four different models were used to estimate SF_ThS-induced fluxes of potential temperature, , heat, , salinity, , and buoyancy, along with potential temperature and salinity diffusivities, and . These were: i) the parametrization of
Table 1
| Parameter symbol | Parameter name |
|---|---|
| Salinity, potential temperature, temperature, heat | |
| Vertical, salt fingering - diffusivity rates of salinity and potential temperature. | |
| Molecular diffusivity rate of temperature | |
| Vertical, salt fingering - fluxes, of salinity, potential temperature and heat. | |
| Vertical, salt fingering - fluxes of salinity-induced and of potential-temperature induced buoyancies, and total, vertical, salt fingering - flux of buoyancy, | |
| = | Density ratio, the ratio of the potential temperature-induced component of density variation with depth over the respective salinity-induced component |
| Flux ratio | |
| Constant of 4/3 power law for buoyancy flux | |
| Coefficient of heat expansion of seawater | |
| Coefficient of saline contraction of seawater | |
| Gravity acceleration | |
| Specific heat capacity of seawater |
Terminology.
Table 2
| Model | Primary formulas | Secondarily derived |
|---|---|---|
| RS12 | , , , , , , | |
| L94 | , | , , , , , , |
| Z98 | , | , , , , , , |
| LAB-4/3 | , | , , , , , |
| Secondary derivation formulas in terms of the primary formulas | ||
| , , , , , , , |
Formulation of the SF_ThS Flux models of Radko and Smith 2012, (RS12),
The application of the semi-empirical models of L94, Z98 started off with calculation, per interface, of salt diffusivities , in terms of the background density ratio (a different formula per model). Subsequently, potential temperature diffusivities, could be determined in terms of and the flux ratio γ, which was taken constant and equal to 0.7, i.e. . Thereafter, in sequence, the fluxes of salt, , potential temperature, heat content, , and salt and heat components of buoyancy fluxes, and along with their resultant, total buoyancy fluxes , were calculated in terms of , and the interfacial, vertical gradients , of potential temperature and of salinity, as follows: ; ; , being the seawater density and the specific heat capacity; ; , and respectively being the coefficients of saline contraction and heat expansion of seawater, and the gravitational constant, and ultimately, . The application of the theoretical model RS12, followed the same methodological route as the afore-mentioned ones of L14 and Z98. The distinction in RS12 considered here was the calculation of in terms of , instead of assigning, as in L14 and Z98, the constant flux ratio value, =0.7. Finally, the use of model LAB_4/3 was initiated by the estimation of the total buoyancy flux , in terms of the interfacial salinity increase, . Thereupon, by using just as for L14, Z98, a constant value of flux ratio =0.7, we applied the following formulas in sequence: ; ; ; ; and finally, . Tables 1, 2, summarize the terminology and formulation of the four models.
The calculations derived smoothed out flux profiles of Potential Temperature (, Salinity, , Heat, (, and Buoyancy, ( versus depth . Fifty percent–overlapping sliding window means were implemented, using a width of 2 data points for the “thick interface” methods of Large (1994, L94), Zhang et al. (1998), Z98), and
3 Results
3.1 Monitoring the SF_ThS
The March 2023 CTD profile in the eastern Cretan Sea (red dot in Figure 1) revealed the existence of a well-defined SF_ThS (Figure 2). The background (or bulk) stability ratio, , calculated by linear regression on the data-curve within the depth-range between the local salinity maximum in the intermediate layer (250 dbar) and the local salinity minimum of the TMW core (1090 dbar) amounts to 1.53. If, on the other hand, the local salinity maximum at 546 dbar is used instead of the intermediate one at 250 dbar, then drops to 1.30. It should be noted here that in contrast to other Mediterranean regions where an intermediate salinity maximum is clearly discernible, this is not the case in the Cretan basin which is a source of intermediate water. The latter value of , is quite close to the one of 1.28 estimated by
Table 3
| L# | Pressure db | Lw db | ° C | psu | m | ° C | ΔS psu | Rρ | Tu | |
|---|---|---|---|---|---|---|---|---|---|---|
| Top | Bot | |||||||||
| 1 | 746 | 766 | 20 | 14.609 | 39.047 | |||||
| 10 | 0.068 | 0.018 | 1.18 | 85.4 | ||||||
| 2 | 776 | 804 | 28 | 14.543 | 39.029 | |||||
| 3 | 0.061 | 0.016 | 1.21 | 84.7 | ||||||
| 3 | 807 | 837 | 30 | 14.477 | 39.012 | |||||
| 10 | 0.070 | 0.018 | 1.18 | 85.3 | ||||||
| 4 | 847 | 882 | 35 | 14.408 | 38.995 | |||||
| 6 | 0.043 | 0.011 | 1.20 | 84.8 | ||||||
| 5 | 888 | 901 | 13 | 14.363 | 38.983 | |||||
Characteristics of the observed staircase in the Cretan Sea: layer sequence number (L#), pressure of layer top and bottom, layer width (Lw), and layer-averaged potential temperature and salinity , interfacial height, , trans-interfacial potential temperature and salinity differences, and ΔS, interfacial stability ratio and angle Turner, Rρ and Tu.
Rρ values have been calculated by a linear fit of the interfacial and values.
During the same month (March 2023), a SeaExplorer glider mission of HCMR sampled along a zonal track across the Cretan Sea, producing a total of 54 deep (down to 970 dbar) temperature and salinity profiles along this track. Thermohaline steps were observed only on the easternmost glider-CTD profile, located in close vicinity to the shipborne CTD profile with the SF_ThS. These profiles were sampled only two days apart (March 17 and March 19, respectively) and were separated by 9.2 km. As shown in Figure 3, only two steps are clearly identifiable in the glider-CTD profile. The first step appears between 734 and 753 dbar, followed by a 10 dbar interface, while the second step lies between 763 and 779 dbar.
Figure 3

SeaExplorer glider cast of March 19th, 2023. Cast position: latitude 35° 46.47 N, longitude 026° 07.82 E. Potential temperature (theta) and salinity profile in the 650–970 dbar layer.
3.2 Fluxes and diffusivities of the SF_ThS. Comparisons with other studies
As visualized in Figure 1 of
Alternatively, the substitution of the previously mentioned bulk stability ratio over the depth regime (547 to 1090 dbar), which apart from encompassing the SF_ThS, extends further up/down to the two mass cores (LIW/CIW and TMW), has yielded bulk fluxes such as and subsequently bulk diffusivities, such as , wherein and accounts for the fact that the mixing-effective portion of the total bulk-salinity gradient, , lies only within the four interfaces of height rather than within the whole staircase regime of depth , as suggested by
The calculations of the flux profiles are tabulated in Table 4 and visualized in Figure 4, while the diffusivity calculations are tabulated/visualized in Table 5 and Figure 5 respectively. One can confirm in the latter figures that the bulk and interfacial values of both fluxes and diffusivities are in relative agreement. The three models that are dependent on the stability ratio/gradients of the interfaces of the SF_ThS, yield diffusivity profiles (e.g., for salinity) that are relatively constant with depth, in contrast to the flux profiles which on average exhibit a tendency of weakening downward fluxes with depth, especially when the latter fluxes are smoothed by a two-point running mean. Particularly, the latter convergence of the smoothed flux-profiles is largely shaped by the influence of the positive flux overshoot at the second interface of the depth regime (804 - 807 dbar) on the running mean. For example, focusing on the salinity fluxes, of the latter interface is much larger than , and as can be clearly seen in Table 5. This occurs because is much larger than for the other interfaces. Indeed, the second interface has a height 2 – 3 times smaller (3 m, compared with 10 m, 10 m, and 6 m for the other three interfaces), while exhibiting comparable across-interface property variation ( for salinity). This gradient sharpening is crucial for enhancing mixing via the downward transport of the salinity surplus and its respective flux, as emphasized by
Table 4
| I # | Pressure (dbar) | Rp | Ks | KΘ | Ks | KΘ | Ks | KΘ | Ks | KΘ | |
|---|---|---|---|---|---|---|---|---|---|---|---|
| ×10–4 m2sec-1 | |||||||||||
| Top | Bott | L94 | Z98 | RS12 | LAB_4/3 | ||||||
| 1 | 766 | 776 | 1.18 | 9.7 | 6.23 | 9.1 | 5.87 | 0.6 | 0.38 | 0.10–0.27 | 0.07–0.17 |
| 2 | 804 | 807 | 1.21 | 8 | 4.51 | 8.26 | 4.66 | 0.37 | 0.19 | 0.28 | 0.15 |
| 3 | 837 | 847 | 1.18 | 9.48 | 5.92 | 9 | 5.62 | 0.52 | 0.31 | 0.22–0.55 | 0.12–0.28 |
| 4 | 882 | 888 | 1.20 | 8.96 | 5.36 | 8.72 | 5.21 | 0.44 | 0.24 | 0.07 | 0.03 |
| stair | 766 | 888 | 1.30 | 8.62 | 5.05 | 8.56 | 5.0 | 0.41 | 0.21 | ||
| avg | 1.19 | 9 | 5.5 | 8.8 | 5.3 | 0.41 | 0.28 | 0.24 | 0.15 | ||
| stder | 0.08 | 0.4 | 0.4 | 0.2 | 0.3 | 0.03 | 0.02 | 0.13 | 0.07 | ||
Vertical salt and heat diffusivities , , over interfaces 1 to 4 and over the whole staircase, along with average interfacial (avg) diffusivities with 95% confidence interval (stder).
Figure 4

Panels (a–d) show for each of the four models (
Table 5
| I # | (×10–6 K m/s) | (×10–6 psu m/s) | (×10–9 W/kg) | (Wm-2) | |
|---|---|---|---|---|---|
| RS12 | 1 | 0.23 | 0.10 | 0.23 | 0.91 |
| 2 | 0.74 | 0.36 | 0.96 | 3.01 | |
| 3 | 0.2 | 0.09 | 0.22 | 0.83 | |
| 4 | 0.17 | 0.08 | 0.20 | 0.69 | |
| stair | 0.15 | 0.07 | 0.18 | 0.61 | |
| L94 | 1 | 3.73 | 1.63 | 3.563 | 15.17 |
| 2 | 17.7 | 7.71 | 16.89 | 71.88 | |
| 3 | 3.9 | 1.70 | 3.724 | 15.85 | |
| 4 | 3.75 | 1.64 | 3.582 | 15.24 | |
| stair | 3.43 | 1.50 | 3.279 | 13.98 | |
| LAB_4/3 | 1 | 0.24-0.45 | 0.13-0.19 | 0.23-0.43 | 0.99-1.82 |
| 2 | 1.3 | 0.55 | 0.12 | 5.18 | |
| 3 | 0.18-0.42 | 0.08-0.17 | 0.17-0.41 | 0.74-1.72 | |
| 4 | 0.04 | 0.02 | 0.04 | 0.16 | |
| stair | – | – | – | – | |
| Z98 | 1 | 3.621 | 1.58 | 3.429 | 14.74 |
| 2 | 19.38 | 8.42 | 18.27 | 78.86 | |
| 3 | 3.834 | 1.67 | 3.625 | 15.60 | |
| 4 | 3.811 | 1.66 | 3.61 | 15.50 | |
| stair | 3.577 | 1.56 | 3.374 | 14.58 |
Downward fluxes of potential temperature, salt, buoyancy, and heat () in the interfaces.
is calculated with from TEOS-10 thermodynamic equation of seawater (IOC, SCOR, & IAPSO, 2010) and the ‘stair’ values correspond to the ‘bulk’ estimations over the whole staircase, e.g. explained in section 4.2. For the LAB_4/3 method, we use a range of values for each of the four “major” interfaces, since this method provides a value for each of the “minor” interfaces within them.
Figure 5

Interfacial diffusivity profile predictions (small symbols) of the four models (RS12, L14, LAB_4/3 and Z98), for potential temperature (a) and salinity (b). The large symbols represent the single, per-model predictions over the whole SF_ThS for three of the four models (excluding the LAB_4/3).
3.3 The susceptibility of the Cretan Sea to salt fingering and SF_ThSs
Using the whole glider dataset, the Turner angle was calculated for each sampling depth in the layer between 250 dbar and the end of each cast. The ceiling of this layer is taken as a typical LIW depth inside the Cretan Sea, while the layer floor corresponds to the maximum glider deployment depth of 970 dbar, which is as close as possible to the TMW core. A histogram showing the normalized distribution of the Turner angle from the glider dataset is presented in Figure 6.
Figure 6

Normalized distribution of the Turner angle in the 250-dbar to end-of-cast layer from the March 2023 HCMR glider dataset. The histogram is divided into 15° segments. Salt fingering, diffusive convection, stable conditions, and unstable conditions are shown in the inset color legend. The inset map shows the position of the glider profiles. The red arrow indicates the position of the shipborne CTD cast shown in Figure 2 relative to the glider profiles.
To investigate the hydrological status of the Cretan Sea during the last decade with respect to the basin’s proneness to double diffusion, shipborne CTD profiles from three different cruises (2016, 2019, and 2021) in the Cretan Sea were examined. These cruises were selected because they collected data along a west-to-east transect, covering most of the open Cretan Sea longitude. For each cruise, we calculated the Turner angle at each sampling depth in the layer between 250 dbar and the deep salinity minimum corresponding to the TMW core. In the EMed, where the upper layers are mostly covered by very saline waters (
Figure 7

(a) Sampling sites of the CRELEV, PERLE 2, PERLE 4, and glider cruises in the Cretan Sea. The observed salinity staircase site shown in Figure 1 is also indicated. (b) Salinity profiles of the CRELEV, PERLE 2, PERLE 4, and glider cruises in the Cretan Sea. Data between 100 and 1,100 dbar are shown. Red lines denote the mean TMW core depth (local salinity minimum) for each cruise. For the glider data, profiles are restricted to 970 dbar, the maximum deployment depth of the glider. This was shallower than the TMW core, and therefore the local salinity minimum was not reached.
Figure 8

Normalized distribution of the Turner angle from 250 dbar to the deep salinity minimum corresponding to the mean TMW core depth for the following: CRELEV 2016 cruise (a), PERLE 2 2019 cruise (b), and PERLE 4 2021 cruise (c). The histogram is divided into 15° segments. For color coding, refer to the inset legend.
4 Discussion and conclusions
As already mentioned, the water mass distribution in the Cretan Sea—i.e., warm, saline LIW/CIW overlying cold, less saline TMW—favors salt fingering processes at the interfaces between these water masses. This is supported by the analysis of Turner angle distributions of in situ data presented here, where cruise data spanning 2016–2023 show that more than 70% of the layer between the intermediate water and the TMW core is prone to salt fingering. However, the occurrence of a thermohaline staircase in the Cretan Sea is rarely reported, previously documented only once (
Overall, LAB_4/3 predicted fluxes and diffusivities similar to those of RS12. The average , values for the Cretan Sea in 2012–2013, (see in
Additionally, the buoyancy fluxes predicted by the models ranged from 10-10W/kg (RS12 and LAB_4/3) to 10-8 W/kg (L14 and Z98). Our LAB_4/3 predictions are one order of magnitude smaller than those obtained in the framework of a major SF-TS monitoring experiment in the western tropical North Atlantic, the C-SALT project (
In any case, SF-TS occurrences are important for the hydrology and climatic variability of the Cretan Sea. As salt fingering transports salt toward deeper layers, this double diffusive process can contribute to the gradual erosion—at least of the upper part—of the TMW low-salinity core within the Cretan Sea. This process drives the core to greater depths with increasing salinity values, as reported by
Moreover, the upward buoyancy flux induced by the vertical motions of the fingers increases stratification, which in turn impedes dense water formation. This hindrance may reduce the role of the Cretan Sea as one of the most important dense water sources of the EMed, despite the invariable winter heat loss observed over the basin since 1950 (
An additional effect of salt fingering in the Cretan Sea concerns nutrient supply to the intermediate water. In particular, SF-DDCs induce a net upward nutrient flux, thereby steepening the positive (increasing with depth) vertical nutrient gradient recorded by
In conclusion, SF-DDCs in the Cretan Sea have not yet been adequately monitored and analyzed, although the basin’s water mass distribution favors their ubiquitous presence. Future work should therefore focus on further investigating these processes, particularly their most pronounced manifestations in the form of SF-TSs, since they are linked to important marine ecosystem functions and respond to pressures induced by climate change.
Statements
Data availability statement
The raw data supporting the conclusions of this article will be made available by the authors, without undue reservation.
Author contributions
DV: Visualization, Formal analysis, Data curation, Writing – original draft, Conceptualization, Methodology, Writing – review & editing, Investigation. SK: Visualization, Formal analysis, Writing – original draft, Data curation, Conceptualization, Methodology, Writing – review & editing, Investigation. EB: Investigation, Data curation, Writing – original draft. DB: Data curation, Investigation, Writing – original draft. VZ: Writing – review & editing.
Funding
The author(s) declare financial support was received for the research and/or publication of this article. The authors acknowledge financial support by the program “Monitoring and recording the status of the marine sub-regions of Greece/Upgrading and functional updating of the MSFD monitoring network”, (MIS 5010880), funded by national Greek and EU funds under the Ministry of Development & Investment, National Strategic Reference Framework (NSRF 2014-2020) and the program “Support for upgrading the operation of the National Network for Climate Change - CLIMPACT” (Grant Agreement: 2023NA11900001, MIS 5201588), funded by national Greek funds under the Ministry of Development & Investment.
Acknowledgments
The authors would like to thank the officers, crews and scientific personnel of the research vessels that collected the data used in this work.
Conflict of interest
The authors declare that the research was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest.
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Supplementary material
The Supplementary Material for this article can be found online at: https://www.frontiersin.org/articles/10.3389/fmars.2025.1649311/full#supplementary-material
References
1
BoscoloR.BrydenH. (2001). Causes of long-term changes in aegean sea deep water. Oceanol. Acta24, 519–5 27. doi: 10.1016/S0399-1784(01)01172-0
2
BrydenH. L.SchroederK.SparnocchiaS.BorghiniM.VetranoA. (2014). Thermohaline staircases in the western Mediterranean sea. J. Mar. Res.72, 1–18. doi: 10.1357/002224014812655198
3
BuffettG. G.KrahmannG.KlaeschenD.SchroederK.SallarèsV.PapenbergC.et al. (2017). Seismic oceanography in the Tyrrhenian Sea: thermohaline staircases, eddies, and internal waves. J. Geophys. Res.122, 8503–8523. doi: 10.1002/2017JC012726
4
ChiggiatoJ.ArtaleV.Durrieu de MadronX.SchroederK.Taupier-LetageI.VelaorasD.et al. (2023). “Recent changes in the mediterranean sea,” in Oceanography of the mediterranean sea. Eds. SchroederK.ChiggiatoJ. (Elsevier), 289–334. doi: 10.1016/B978-0-12-823692-5.00008-X
5
ConanP.Pujo-PayM.Durrieu de MadronX. (2019). PERLE. doi: 10.18142/348
6
DuranteS.OliveriP.NairR.SparnocchiaS. (2021). Mixing in the tyrrhenian interior due to thermohaline staircases. Front. Mar. Sci.8. doi: 10.3389/fmars.2021.672437
7
DuranteS.SchroederK.MazzeiL.PieriniS.BorghiniM.SparnocchiaS. (2019). Permanent thermohaline staircases in the Tyrrhenian Sea. Geophys. Res. Lett.46, 1562–1570. doi: 10.1029/2018GL081747
8
Durrieu de MadronX.BlinP.Pujo-PayM.TaillandierV.ConanP. (2024). Effect of double diffusion processes in the deep ocean on the distribution and dynamics of particulate and dissolved matter: a case study in Tyrrhenian Sea, EGUsphere [preprint]. doi: 10.5194/egusphere-2024-3436
9
EMODnet Bathymetry Consortium (2016). “EMODnet digital bathymetry (DTM 2016),” in EMODnet bathymetry consortium. doi: 10.12770/c7b53704-999d-4721-b1a3-04ec60c87238
10
FalcoP.TraniM.ZambianchiE. (2016). Water mass structure and deep mixing processes in the Tyrrhenian Sea: Results from the VECTOR project. Deep-Sea Res. I: Oceanogr. Res. Pap.113, 7–21. doi: 10.1016/j.dsr.2016.04.002
11
FerronB.Bouruet-AubertotP.SchroederK.BrydenH. L.CuypersY.BorghiniM. (2021). Contribution of thermohaline staircases to deep water mass modifications in the western mediterranean sea from microstructure observations. Front. Mar. Sci.8. doi: 10.3389/fmars.2021.664509
12
GlessmerM. S.OschliesA.YoolA. (2008). Simulated impact of double-diffusive mixing on physical and biogeochemical upper ocean properties. J. Geophys. Res. Oceans113, C08029. doi: 10.1029/2007JC004455
13
GreggM. C.AlfordM. H.KontoyiannisH.ZervakisV.WinkelD. (2012). Mixing over the steep side of the Cycladic Plateau in the Aegean Sea. J. Mar. Sys.89, 30–47. doi: 10.1016/j.jmarsys.2011.07.009
14
GregM. C.SanfordT. B. (1987). Shear and turbulence in thermohaline staircases. Deep-Sea Res. A. Oceanogr. Res. Pap.24 (10), 1689–1696. doi: 10.1016/0198-0149(87)90017-3
15
IOCSCORIAPSO (2010). The international thermodynamic equation of seawater–2010: Calculation and use of thermodynamic properties. Intergovernmental Oceanographic Commission, Manuals and Guides 56, UNESCO(English), 196pp.
16
JohnsonG. C.KearneyK. A. (2009). Ocean climate change fingerprints attenuated by salt fingering? Geophys. Res. Lett.36, L21603. doi: 10.1029/2009GL040697
17
JoseyS. A.SchroederK. (2023). Declining winter heat loss threatens continuing ocean convection at a Mediterranean dense water formation site. Environ. Res. Lett.18, 024005. doi: 10.1088/1748-9326/aca9e4
18
KelleyD. E. (1984). Effective diffusivities within oceanic thermohaline staircases. J. Geophys. Res.89, 10484–10488. doi: 10.1029/JC089iC06p10484
19
KelleyD. E. (1990). Fluxes through diffusive staircases: a new formulation. J. Geophys. Res.95, 3365–3371. doi: 10.1029/JC095iC03p03365
20
KelleyD. E.FernandoH. J. S.GargettA. E.TannyJ.ÖzsoyE. (2003). The diffusive regime of double-diffusive convection. Prog. Oceanogr.56, 461–481. doi: 10.1016/S0079-6611(03)00026-0
21
KhodayarS.PastorF.ValienteJ. A.BenetóP.EhmeleF. (2022). What causes a heavy precipitation period to become extreme? The exceptional October of 2018 in the Western Mediterranean. Weather Clim. Extrem.38, 100493. doi: 10.1016/j.wace.2022.100493
22
KilicarslanB. M.YucelI.PilatinH.DuzenliE.YilmazM. T. (2021). Improving WRF-Hydro runoff simulations of heavy floods through the sea surface temperature fields with higher spatio-temporal resolution. Hydrol. Process.35, e14338. doi: 10.1002/hyp.14338
23
KioroglouS.TragouE.ZervakisV.GeorgopoulosD.HerutB.GertmanI.et al. (2014). Vertical diffusion processes in the Eastern Mediterranean — Black Sea System. J. Mar. Sys., 135, 53–63. doi: 10.1016/j.jmarsys.2013.08.007
24
KokoszkaF.IudiconeD.ZingoneA.SaggiomoV.Ribera D’AlcaláM.ConversanoF. (2021). A note about density staircases in the Gulf of Naples: 20 years of persistent weak salt-fingering layers in a coastal area. Adv. Oceanogr. Limnol.12. doi: 10.4081/aiol.2021.10008
25
KontoyiannisH.LykousisV.PapadopoulosV.StavrakakisS.AnassontzisE. G.BeliasA.et al. (2016). Hydrography, Circulation, and Mixing at the Calypso Deep (the Deepest Mediterranean Trough) during 2006 – 09. J. Phys. Oceanogr.46, 1255–1276. doi: 10.1175/JPO-D-15-0198.1
26
KubinE.PoulainP.-M.MauriE.MennaM.NotarstefanoG. (2019). Levantine intermediate and levantine deep water formation: an argo float study from 2001 to 2017. Water11, 1781. doi: 10.3390/w11091781
27
KunzeE. (1987). Limits on growing, finite-length salt fingers: A Richardson number constraint. J. Mar. Res.45, 533–566. doi: 10.1357/002224087788326885
28
LargeW. G.McWilliamsJ. C.DoneyS. C. (1994). Oceanic vertical mixing: A review and a model with a nonlocal boundary layer parameterization. Rev. Geophys.32, 363–403. doi: 10.1029/94RG01872
29
LarsonN. G.GreggM. C. (1983). Turbulent dissipation and shear in thermohaline intrusions. Nature306, 26–32. doi: 10.1038/306026a0
30
LindenP. F. (1973). On the structure of salt fingers. Deep-Sea Res.20, 325–340. doi: 10.1016/0011-7471(73)90057-0
31
LIWEX Group (2003). The Levantine Intermediate Water Experiment (LIWEX) Group: Levantine basin—A laboratory for multiple water mass formation processes. J. Geophys. Res.108, 8101. doi: 10.1029/2002JC001643
32
LueckR. G. (1987). Microstructure measurements in a thcrmohaline staircase. Deep-Sea Res.34, 1677–1688. doi: 10.1016/0198-0149(87)90016-1
33
MarmorinoG. O.BrownW. K.MorrisW. D. (1987). Two-dimensional temperature structure in the C-SALT thermohaline staircase. Deep-Sea Res.34, 1667–1676. doi: 10.1016/0198-0149(87)90015-X
34
MartínezJ.LeonelliF. E.García-LadonaE.GarrabouJ.KerstingD. K.BensoussanN.et al. (2023). Evolution of marine heatwaves in warming seas: the Mediterranean Sea case study. Front. Mar. Sci.10. doi: 10.3389/fmars.2023.1193164
35
McDougallT. J.TaylorJ. R. (1984). Flux measurements across a finger interface at low values of the stability ratio. J. Mar. Res.42, 1–14. doi: 10.1357/002224084788506095
36
MecciaV. L.SimoncelliS.SparnocchiaS. (2016). Decadal variability of the Turner Angle in the Mediterranean Sea and its implications for double diffusion. Deep-Sea Res. I: Oceanogr. Res. Pap.114, 64–77. doi: 10.1016/j.dsr.2016.04.001
37
OschliesA.DietzeH.KählerP. (2003). Salt-finger driven enhancement of upper ocean nutrient supply. Geophys. Res. Lett.30, 3–6. doi: 10.1029/2003GL018552
38
OzerT.GertmanI.KressN.SilvermanJ.HerutB. (2017). Interannual thermohaline, (1979 – 2014) and nutrient, (2002 – 2014) dynamics in the Levantine surface and intermediate water masses, SE Mediterranean Sea. Glob. Plan. Change151, 60–67. doi: 10.1016/j.gloplacha.2016.04.001
39
PavlidouA.VelaorasD.KarageorgisA. P.RousselakiE.ParinosC.DähnkeK.et al. (2020). Seasonal variations of biochemical and optical properties, physical dynamics and N sta ble isotopic composition in three northeastern Mediterranean basins (Aegean, Cretan and Ionian Seas). Deep-Sea Res. II: Top. Stud. Oceanogr.171, 104704. doi: 10.1016/j.dsr2.2019.104704
40
POEM Group (1992). General circulation of the eastern mediterranean. Earth-Sci. Rev.32, 285–309. doi: 10.1016/0012-8252(92)90002-B
41
PotirisM.MamoutosI. G.TragouE.ZervakisV.KassisD.BallasD. (2024). Dense water formation variability in the aegean sea from 1947 to 2023. Oceans5, 611–6 36. doi: 10.3390/oceans5030035
42
RadcoT.SmithD. P. (2012). Equilibrium transport in double-diffusive convection. J. Fluid Mech.692, 5–27. doi: 10.1017/jfm.2011.343
43
RadkoT. (2013). Double-diffusive convection (Cambridge: Cambridge University Press), 344. doi: 10.1017/CBO9781139034173
44
RuddickB. (1983). A practical indicator of the stability of the water column to double-diffusive activity. Deep-Sea Res. A. Oceanogr. Res. Pap.30, 1105–1107. doi: 10.1016/0198-0149(83)90063-8
45
SchmittR. W. (1979). Flux measurements at an interface. J. Mar. Res.37, 419–436.
46
SchmittR. W.PerkinsH.BoydJ. D.StalcupM. C. (1987). C-SALT: An investigation of the thermohaline staircase in the western tropical North Atlantic. Deep Sea Res.34, 1655–1665. doi: 10.1016/0198-0149(87)90014-8
47
SchroederK.JoseyS. A.HerrmannM.GrignonL.GaspariniG. P.BrydenH. L. (2010). Abrupt warming and salting of the Western Mediterranean Deep Water after 2005: Atmospheric forcings and lateral advection. J. Geophys. Res. Oceans115, 1–18. doi: 10.1029/2009JC005749
48
SternM. E. (1960). The “Salt-fountain“ and thermohaline convection. Tellus12, 172–175. doi: 10.3402/tellusa.v12i2.9378
49
SternM. ,. E. (1976). Maximum buoyancy flux across a salt finger interface. J. Mar. Res.34, 95–11.
50
TaillandierV.D’OrtenzioF.PrieurL.ConanP.CoppolaL.CornecM.et al. (2022). Sources of the Levantine intermediate water in winter 2019. J. @ Geophys. Res. Oceans127, e2021JC017506. doi: 10.1029/2021JC017506
51
TaillandierV.PrieurL.D’OrtenzioF.Ribera d’AlcalàM.Pulido-VillenaE. (2020). Profiling float observation of thermohaline staircases in the western Mediterranean Sea and impact on nutrient fluxes. Biogeosciences17, 3343–3366. doi: 10.5194/bg-17-3343-2020
52
TennekesH.LumleyJ. L. (1972). A first course in turbulence (Cambridge, MA: The MIT Press), 293.
53
TheocharisA.GeorgopoulosD.LascaratosA.NittisK. (1993). Water masses and circulation in the central region of the Eastern Mediterranean. Deep-Sea Res. II40, 1121–1142. doi: 10.1016/0967-0645(93)90064-T
54
TheocharisA.KleinB.NittisK.RoetherW. (2002). Evolution and status of the eastern mediterranean transien- 1999). J. Mar. Sys.33 – 34, 91–116. doi: 10.1016/S0924-7963(02)00054-4
55
TheocharisA.NittisK.KontoyiannisH.PapageorgiouE.BalopoulosE. (1999). Climatic changes in the Aegean Sea influence the eastern Mediterranean thermocline circulatio- 1997). Geophys. Res. Lett.26, 1617–1620. doi: 10.1029/1999GL900320
56
TurnerJ. S. (1967). Salt fingers across a density interface. Deep-Sea Res.14, 599–611. doi: 10.1016/0011-7471(67)90066-6
57
VelaorasD.KrokosG.NittisK.TheocharisA. (2014). Dense intermediate water outflow from the Cretan Sea: A salinity driven, recurrent phenomenon, connected to thermohaline circulation changes. J. Geophys. Res. Oceans119, 4797–4820. doi: 10.1002/2014JC009937
58
VelaorasD.KrokosG.TheocharisA. (2015). Recurrent intrusions of transitional waters of Eastern Mediterranean origin in the Cretan Sea as a tracer of Aegean Sea dense water formation events. Progr. Oceanogr.135, 113–124. doi: 10.1016/j.pocean.2015.04.010
59
VelaorasD.PapadopoulosV. P.KontoyiannisH.CardinV.CivitareseG. (2019). Water masses and hydrography during April and June 2016 in the Cretan Sea and Cretan Passage (Eastern Mediterranean Sea). Deep-Sea Res. II: Top. Stud. Oceanogr.164, 25–40. doi: 10.1016/j.dsr2.2018.09.005
60
VelaorasD.ZervakisV.TheocharisA. (2021). The physical characteristics and dynamics of the aegean water masses (Berlin Heidelberg: Springer), 1–29. doi: 10.1007/698_2020_730
61
YouY. (2002). A global ocean climatological atlas of the Turner angle: Implications for double-diffusion and water-mass structure. Deep-Sea Res. Part I: Oceanogr. Res. Pap.49, 2075–2093. doi: 10.1016/S0967-0637(02)00099-7
62
ZhangJ.SchmittR. W.HuangR. X. (1998). Sensitivity of the GFDL modular ocean model to parameterization of double-diffusive processes. J. Phys. Oceanogr.28, 589–605. doi: 10.1175/1520-0485(1998)028<0589:SOTGMO>2.0.CO;2
63
ZodiatisG. (1990). Double Diffusive Activities in the Cretan Sea during late summer 1987. Rapp. Commun. Int. Mer Medit.32, 1, 189.
64
ZodiatisG. (1991). The multi-scaled thermohaline structure of the cretan sea waters, ph.D. Thesis Vol. 551.465 (St. Petersburg, UDK: LHMI (RSHM University), 191.
65
ZodiatisG. (1992). Lens formation in the SE Ionian Sea and double diffusion. Annales Geophysicae10, 935–941.
66
ZodiatisG.GaspariniG. P. (1996). Thermohaline staircase formations in the Tyrrhenian Sea. Deep-Sea Res. I: Oceanogr. Res. Pap.43, 655–678. doi: 10.1016/0967-0637(96)00032-5
Summary
Keywords
Eastern Mediterranean Sea, Cretan Sea, double diffusive convection, salt fingering, thermohaline staircase, heat and salt diffusivities, heat and salt fluxes
Citation
Velaoras D, Kioroglou S, Bourma E, Ballas D and Zervakis V (2025) Evidence of thermohaline staircase in the Cretan Sea (Eastern Mediterranean Sea). Front. Mar. Sci. 12:1649311. doi: 10.3389/fmars.2025.1649311
Received
18 June 2025
Accepted
11 August 2025
Published
09 September 2025
Volume
12 - 2025
Edited by
Donglai Gong, College of William & Mary, United States
Reviewed by
Sara Durante, National Research Council (CNR), Italy
Anıl Akpınar, Middle East Technical University, Türkiye
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© 2025 Velaoras, Kioroglou, Bourma, Ballas and Zervakis.
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*Correspondence: Dimitris Velaoras, dvelaoras@hcmr.gr
†These authors have contributed equally to this work
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