REVIEW article

Front. Neurol., 08 March 2019

Sec. Movement Disorders

Volume 10 - 2019 | https://doi.org/10.3389/fneur.2019.00195

Biomarkers for Dementia, Fatigue, and Depression in Parkinson's Disease

  • 1. Department of Neurology, Jena University Hospital, Jena, Germany

  • 2. Center for Healthy Ageing, Jena University Hospital, Jena, Germany

Abstract

Parkinson's disease is a common multisystem neurodegenerative disorder characterized by typical motor and non-motor symptoms. There is an urgent need for biomarkers for assessment of disease severity, complications and prognosis. In addition, biomarkers reporting the underlying pathophysiology assist in understanding the disease and developing neuroprotective therapies. Ultimately, biomarkers could be used to develop a more efficient personalized approach for clinical trials and treatment strategies. With the goal to improve quality of life in Parkinson's disease it is essential to understand and objectively monitor non-motor symptoms. This narrative review provides an overview of recent developments of biomarkers (biofluid samples and imaging) for three common neuropsychological syndromes in Parkinson's disease: dementia, fatigue, and depression.

Introduction

Parkinson's disease (PD) is now considered as progressive and multisystem α-synucleinopathy. Therefore, PD is characterized not only by motor symptoms, but also a broad range of non-motor symptoms (NMS) (1). NMS can aggravate disease burden and significantly contribute to worsening of quality of life (2). Biomarkers which are associated with worse motor performance as well as development of NMS are of special importance in PD. A biomarker is “a characteristic that is objectively measured and evaluated as an indicator of normal biological processes, pathogenic processes, or pharmacologic responses to a therapeutic intervention” (3). The ideal PD biomarkers should have a reasonable effect size, are reproducible across different cohorts and are ideally verified in neuropathological proven PD cases. Biomarkers in PD can include (i) biomarker for prodromal stage to identify PD before motor symptoms occur, (ii) biomarkers of susceptibility to identify persons who are at risk for PD, (iii) biomarkers for motor and non-motor burden to assess disease severity and monitor the efficacy of therapies. The last one can help to identify patients who are at risk to develop complications and may lead to individual optimization and prevention in health care. This review provides an update on recent advances in the development of biomarkers (biofluid samples and neuroimaging) for three common neuropsychological syndromes: dementia, fatigue and depression.

Cognitive Impairment

Cognitive deficits are common in PD and can present as mild dysfunction in the prodromal and early stages, or as dementia (PDD) in advanced stages (4). Approximately 20% of patients with de novo PD have mild cognitive impairment (MCI) (5). The concept of PD-MCI was introduced 2012 (MDS Task Force) and characterizes a cognitive decline that is assessed during neuropsychological testing but does not impair activities of daily living (6). MCI is considered an intermediate state of cognitive dysfunction in PD that may progress to PDD. Up to 75% of patients will develop dementia over the longterm disease course (7). However, the rate to PDD, the cognitive profile and severity of cognitive dysfunction show high interindividual variation. Given its high medical and social impact and its health-related costs, the identification of biomarkers for PDD is of high priority (8). Biomarkers reflecting cognitive decline can facilitate early diagnosis and may indicate response to therapeutic interventions.

Clinical factors, such as higher age, male sex, low level of education, longer disease duration, higher Hoehn & Yahr stage, axial impairment, excessive daytime sleepiness, cardiovascular autonomic dysfunction, REM sleep behavior disorder, hallucinations and PD-MCI were found to strongly predict the development of PDD (913). Moreover, impairment of memory and language (posterior-cortical dysfunction) seems to be linked to a higher risk of PDD (14, 15).

Given the neuropathology of PDD several studies aimed to identify biomarkers which reflect proteinopathy, neuronal loss, abnormal neurotransmitters, and structural and functional brain changes. Lewy bodies and amyloid plaques in the neocortex and limbic system are typical neuropathological features of Alzheimer's disease and PDD (16, 17). Hence, the majority of studies investigated amyloid-ß 1–42 (Aß), tau protein, and α-synuclein in the cerebrospinal fluid (CSF) of PD patients (Table 1). In many studies the level of Aß was reduced in PDD. Low CSF levels of Aß were found to be related to deterioration in attention, executive function, semantic fluency and memory (21, 38, 40, 45). One-half of PDD patients had the CSF biomarker signature of Alzheimer's disease (46) suggestive of an overlap with Alzheimer's disease pathology (47). Low baseline CSF Aβ was associated with more rapid cognitive decline later in disease. By contrast, the levels of total (t-tau) and phosphorylated tau (p-tau) were found to be increased or unchanged in PDD (Table 1). For clinicians it is highly relevant to know which biomarkers accurately predict the progression from MCI to PDD. Therefore, based on the data from cross-sectional and longitudinal studies one can assume that reduced Aß predicts cognitive decline in PD (40, 42, 48).

Table 1

StudyCSF biomarkerParticipantsMethodsResult
Aß1-42t- taup-taut-α-syno-α-synother
Alves et al. (18)*+++PDND 104MDS Task ForceLow Aβ predicted early dementia
Bäckström et al. (19)*+++++PDND 104
C 30
PSP 13
MSA 11
NFL
H-FABP
Low Aβ, NFL and H-FABP predicted PDD
Brockmann et al. (20)++PDND 353
PDD 103
Genetic variants known to be involved in Aβ clearanceRisk variants in APOE and cystatin C genes were associated with lower Aβ
Compta et al. (21)++PDND 20
PDD 20
C 30
MMSE
DSM-IV-R
MDS Task Force
PDD: ↑ t-tau
PDND: ↓ Aβ positively correlated with phonemic fluency
Compta et al. (22)+++PDND 19
PDD 29
C 9
MMSE
MDS Task Force
PDD: ↓ Aβ
↑ t-tau and p-tau in a subgroup
Compta et al. (23)*+++PDND 27MMSE
MDS Task Force
Low Aβ predicted PDD
Compta et al. (24)++++PDND 21
PDD 20
C 13
MMSE/PDD by MDS Task ForcePDD: ↓ Aβ, ↑ t-tau, ↑ o-α-syn
Ffytche et al. (25)+PD 423
3-4 years of follow-up
Compare baseline structural imaging and CSF data in patients who go on to develop illusions or hallucinations in newly diagnosed PDPatients with early onset PD psychosis: Aβ ↓
Gmitterová et al. (26)++++PDND 22
PDD 31
DLB 51
C 32
Discriminatory potential of tau, p-tau, Aβ, NSE and S100B across the spectrum of LBDPDD Aβ ↓, tau ↑
Rapid disease course not associated with decrease of Aβ
Halbgebauer et al. (27)+PDND 22
PDD 29
C 36
Modified serpinA1PDD: acidic serpinA1 isoform ↑
Hall et al. (28)++++PDND 90
PDD 33
C 107
MMSE
MDS Task Force
PDD: ↑ p-tau, Aβ or t-α-syn no differences
Hall et al. (29)*+++++PDND 42
C 69
Low Aβ predicted memory decline, high α-syn predicted reduced cognitive speed
Hansson et al. (30)++PDND 30
C 98
MMSE
MDS Task Force
PDD: ↑ o-α-syn
Janssens et al. (31)++++probable AD 52
FTD 59
DLB 39
PDD 14
C 88
young C 32
3-methoxy-4-hydroxyphenylglycol (MHPG)Aβ young C > C > FTD > PDD, DLB > AD
tau AD > FTD > PDD, DLB > C > young C
p-tau AD > FTD = PDD,DLB = C> young C
MHPG PDD, DLB > AD > C
Lindqvist et al. (32)+PDND 71
PDD 16
C 33
MMSEPDD: C-reactive protein ↑
IL6 ↑
TNF-Alpha →
Eotaxin →
MCP-1 →
MIP-1beta →
IP-10 →
Maetzler et al. (33)+PDND 14
PDD 12
MMSEPDD: Aβ ↓
Maetzler et al. (34)++PDND 21
PDD 10
C 39
MMSENo difference
Maetzler et al. (35)++PDND 77
PDD 26
C 72
MMSE
MDS Task Force
No difference
Modreanu et al. (36)+++PD 37
PDD 21
PDD at 18-months 35
Spatial disorientation, memory complaints over disease coursePDD: Aβ ↓
tau and p-tau no difference
‘PDD -converters' had significantly lower Aβ at baseline
Parnetti et al. (37)++PDND 67
PDD 48
C 41
MMSENo difference
Parnetti et al. (38)*+++++PDND 44
Disease C 25
MMSE
MoCa
Low Aβ predicted more rapid decline
Schrag et al. (39)*++PDND 390
C 178
MoCa over 2 yearsLow Aβ/t-tau ratio predicts cognitive decline
Siderowf et al. (40)*+++PDND: 45Dementia rating scaleLow Aβ predicted rapid decline in Dementia rating scale
Stewart et al. (41)*++++PDND 350Verbal memory, cognitive processing speed, and visuospatial working memoryLower α-synuclein predicted better preservation of cognitive function
Terrelonge et al. (42)*++++PDND 341Memory, visuospatial, working memory–executive function, and attention processing speedLow Aβ predicted cognitive impairment
Vranová et al. (43)++PDND 27
PDD 14
C 14
MMSE
MDS Task Force
PDD: ↑ t-tau/ Aβ index
Aβ or t-tau no differences
Wennström et al. (44)+
PDND 38
PDD 22
C 52
MMSE
MDS Task Force
No difference

Cerebrospinal-fluid (CSF) biomarkers of cognitive impairment and dementia in Parkinson's disease.

PD, Patients with Parkinson's disease; PD-MCI, Parkinson's disease patients with mild cognitive impairment; PDD, Parkinson's disease patients with dementia; PDND, non-demented PD; MSA, multiple system atrophy; PSP, progressive supranuclear palsy; AD, Patients with Alzheimer's disease; DLB, Dementia with Lewy body; C, Controls; MoCA, Montreal Cognitive Assessment; MMSE, Mini Mental Status Examination; Aβ, Aβ1−42 amyloid; NFL, neurofilament light chain protein; H-FABP, heart fatty acid-binding protein;

*

longitudinal studies.

Several studies assessed the CSF levels of α-synuclein in PD. Meta-analyses demonstrated that total α-synuclein levels are lower in PD compared to controls (49, 50). However, in terms of α-synuclein and cognitive decline there are conflicting results with both low and high levels in the presence of cognitive impairment (29, 41, 48). In the DATATOP study with up to 8 years of follow-up, lower α-synuclein levels predicted better preservation of cognitive function (verbal learning and memory, visuospatial working memory) in early disease. Thus, α-synuclein may reflect changes in multiple cognitive domains and may predict cognitive decline in PD (29, 41, 48). On the other hand most studies of non-demented PD failed to find any association between α-synuclein levels and cognition (51, 52). It seems that CSF α-synuclein levels may increase with disease stage. This could explain why cognitive deficits in connection with high levels of α- synuclein were found in more advanced disease stages (53). Isoforms of α-synuclein (e.g., phosphorylated, ubiquitinated, oligomeric) are potentially more sensitive to cognitive decline than the total α-synuclein level (24, 30). Another study examining plasma levels of α-synuclein found higher levels in PDD and a correlation with mini mental state examination scores (54). This finding, however, requires further investigations.

In another longitudinal study, high neurofilament light chain protein, low Aβ and high heart fatty acid–binding protein at baseline were related to future PDD with a relatively high diagnostic accuracy (19). Also several serum proteins, such as C-reactive protein, interleukins, interferon-γ, tumor necrosis factor α, uric acid, and cystatin C were found to be associated with cognition in PD (55). In particular, low uric acid concentrations, low levels of epidermal growth factor (EGF) and insulin-like growth factor (ILGF) seems to have predictive value for deterioration of cognitive function in PD (5661). In combination with clinical markers, a study of 390 patients from the Progression Markers Initiative study with newly diagnosed PD, the occurrence of cognitive impairment at 2 years follow-up could be predicted with good accuracy using a model combining information on age, non-motor assessments, DAT imaging, and CSF biomarkers. Here, the Montreal Cognitive Assessment (MoCA) scores and low CSF Aβ to t-tau ratio and DAT imaging results were the best predictors of cognitive impairment (39). Using data from the Parkinson's Progression Markers Initiative, Fereshtehnejad et al., identified distinct subgroups via a cluster analysis of a comprehensive dataset consisting of clinical characteristics, neuroimaging, biospecimen and genetic information. Here, the CSF biomarkers differed between these PD subtypes. Patients with diffuse malignant disease course and fast cognitive decline, showed an Alzheimer's disease-like CSF profile (low Aβ, low Aβ/t-tau ratio) (62).

Applying computerized neuroimaging analyses several MRI studies have found gray matter atrophy and disruptions of white matter integrity in PDD, although findings in non-demented PD and PD-MCI remain inconsistent (63) (Tables 2, 3). A longitudinal study using voxel-based morphometry (VBM) found neocortical volume reduction (temporo-occipital region, hippocampal and parahippocampal) as the most relevant finding in patients who develop PDD (97). Another study has identified a validated Alzheimer's disease pattern of brain atrophy as an independent predictor of cognitive impairment in PD (64). More specifically cortical thinning in the right precentral, frontal, and in the anterior cingulate cortex as well as gray matter atrophy (prefrontal, insula, caudate nucleus, hippocampal) predicted cognitive decline in PD (23, 66, 70, 76, 98). Cognitive impairment was also found to be associated with lower gray matter volume and increased mean diffusivity in the nucleus basalis of Meynert, compared to non-demented patients. Moreover, these changes were predictive for developing cognitive impairment in cognitively intact patients with PD, independent of other clinical and non-clinical markers of the disease (99). The nucleus basalis of Meynert and the pedunculopontine nucleus in the brainstem are important cholinergic projections in and post-mortem studies have shown that neuronal loss in in the nucleus basalis is an early phenomenon in PD (100, 101). Combining many modalities, Compta et al. (23) performed a longitudinal study in non-demented PD patients including CSF, neuropsychological and MRI studies at baseline and 18 months follow up. Here, a combination of lower CSF Aβ, reduced verbal learning, semantic fluency and visuoperceptual scores, as well as cortical thinning in superior-frontal/anterior cingulate and precentral regions were found to be predictive for PDD.

Table 2

StudyParticipantsMethodsResult
Weintraub et al. (64)PDND 60VBM*In PD-MCI hippocampal and temporal gray matter atrophy.
Melzer et al. (65)PDND 57
PD-MCI 23
PDD 16
C 34
VBMIn PD-MCI gray matter atrophy in temporal, parietal, frontal cortex, amygdala, right putamen, and hippocampus.
In PDD additional atrophy in medial temporal lobe, lingual gyrus, posterior cingulate gyrus, and bilateral caudate.
Lee et al. (66)PD-MCI 51
C 25
VBM*PD-MCI to PDD converters had lower GM density in the left prefrontal areas, left insular cortex and bilateral caudate nucleus compared with that in PD-MCI non-converters.
Borroni et al. (67)PDND11
PDD 10
LBD 13
C 10
VBMIn PDD bilateral frontal and subcortical (caudate nucleus) gray matter atrophy.
Duncan et al. (68)PDND 125
C 50
VBM
DTI
Frontal and parietal gray matter volume reductions were associated with reduced executive function. Increased mean diffusivity was associated with performance on the semantic fluency and Tower of London tasks in frontal and parietal white matter tracts.
Hattori et al. (69)PDND 32
PD-MCI 28
PDD 25
DLB 29
C 40
VBM
TBSS
In PDD more atrophy in the cerebellum, thalami, insula, parietal cortex and occipital cortex.
Kandiah et al. (70)PDND 97Hippocampal volume
White matter hyperintensities*
Hippocampal volume predicts PD-MCI and PDD.
Rektorova et al. (71)PDND 75
PD-MCI 29
PDD 22
Spatial Independent Component AnalysisIn PDD gray matter volume reductions in the hippocampus and temporal lobes, fronto-parietal regions and increases in the midbrain/cerebellum correlated with visuospatial deficits and letter verbal fluency, respectively.
Biundo et al. (72)PDND 15
PD-MCI 14
HC 21
Cortical thicknessIn PD-MCI cortical thinning in right supramarginal, dorsolateral prefrontal cortex, hippocampus, orbito-frontal, fusiform, superior parietal, and cuneus.
Pereira et al. (73)PDND 90
PD-MCI 33
H 56
Cortical thicknessIn PD-MCI cortical thinning in left precuneus, inferior temporal precentral, superior parietal, and lingual regions.
Hanganu et al. (74)PDND 15
PD-MCI 17
H 18
Cortical thickness *In PD-MCI thinning in temporal and medial occipital lobe, nucleus accumbens and amygdala correlate with cognitive decline.
Ibarretxe-Bilbao et al. (75)PDND 16
C 15
Cortical thickness*In PD cortical thinning in bilateral fronto-temporal regions and reduced amygdala volume.
Mak et al. (76)PDND 66
PD-MCI 39
H 37
Cortical thickness*PD-MCI converters showed bilateral temporal cortex thinning at baseline.
Hwang et al. (77)PDND 12
PDD 11
C 14
Cortical pattern matching
Cortical thickness
PDD showed thinning bilateral sensorimotor, lateral parietal, right posterior cingulate, parieto-occipital, inferior temporal and lateral frontal relative to C and PDND.
Zarei et al. (78)Early PD 24 moderate PD 18
PDD 15
C 39
Cortical thicknessMMSE correlated positively with cortical thickness in the anterior temporal, dorsolateral prefrontal, posterior cingulate, temporal fusiform and occipitotemporal cortex.
Pagonabarraga et al. (79)PDND 26
PD-MCI 26
PDD 20
C 18
Cortical thicknessFrom PDND to PDD a linear and progressive cortical thinning was observed in areas functionally specialized in declarative memory (entorhinal cortex, anterior temporal pole), semantic knowledge (parahippocampus, fusiform gyrus), and visuoperceptive integration (banks of the superior temporal sulcus, lingual gyrus, cuneus and precuneus).
Carlesimo et al. (80)PDND 25
C 25
DTIIncreased mean diffusivity in the PD hippocampi; high hippocampal mean diffusivity values obtained low memory scores.
Chen et al. (81)PDND 19
PDD 11
C 21
DTIIn PDD lower fractional anisotropy in the left hippocampus, higher mean diffusivity in widespread white matter regions. In PD positive correlation between MoCA score and fractional anisotropy of left inferior longitudinal and hippocampus, and bilateral superior longitudinal fasciculus.

Cortical and subcortical structural changes related to cognitive impairment and dementia in Parkinson's disease.

PD, Patients with Parkinson's disease; PD-MCI, Parkinson's disease patients with mild cognitive impairment; PDD, Parkinson's disease patients with dementia; PDND, non-demented PD; DLB, Dementia with Lewy body; C, Controls; MoCA, Montreal Cognitive Assessment; MMSE, Mini Mental Status Examination;

*

longitudinal studies.

Table 3

StudyParticipantsMethodsResult
Gorges et al. (82)PDND 14
PDD 17
C 22
Resting-state fMRIIn PDND hyperconnectivity (network expansions) in cortical, limbic, and basal ganglia-thalamic areas. In PDD decreased intrinsic functional connectivity compared with controls (predominantly between major nodes of the default mode network).
Baggio et al. (83)PDND 32
PD-MCI 23
C 36
Resting-state fMRIIn PD-MCI reduced connectivity between dorsal attention network and right fronto-insular regions (worse performance in executive functions) and increased connectivity between default mode network and medial and lateral occipito-parietal regions (worse visuo-spatial performance).
Amboni et al. (84)PDND 21
PD-MCI 21
C 20
Resting-state fMRIIn PD-MCI patients decreased functional connectivity in bilateral prefrontal cortex (fronto-parietal network).
Tessitore et al. (85)PDNT 16
C 16
Resting-state fMRIIn PDND decreased default mode network connectivity correlated with cognitive parameters.
Rektorova et al. (86)PDND 18
PDD 14
C 18
Resting-state fMRIIn PDD decreased connectivity in the right inferior frontal gyrus compared to PDND and C (using posterior cingulate cortex/precuneus as seed for analysis).
Borroni et al. (67)PDND11
PDD 10
LBD 13
C 10
Resting-state fMRIReduced local coherence of frontal regions in PD and in PDD.
Olde et al. (87)PDND 55
C 15
Resting-state fMRIIn PDND longitudinally decreases in functional connectivity most prominent for posterior brain regions correlated with disease progression and cognitive decline.
Seibert et al. (88)C 19
PDND 19
PDD 18
Resting-state fMRIIn PDD corticostriatal functional correlations were decreased in bilateral prefrontal regions.
Lin et al. (89)PDND 17
PDD 17
C 17
Arterial spin labeling (ASL) magnetic resonance imaging (ASL-MRI)In PDND and PDD progressive widespread cortical hypoperfusion.
Le Heron et al. (90)PDD 20
AD 17
C 37
Arterial spin labeling (ASL) magnetic resonance imaging (ASL-MRI)In AD and PDD posterior hypoperfusion (including posterior cingulate gyrus, precuneus, occipital regions). Perfusion in medial temporal lobes (AD < PDD) and right frontal cortex (PDD < AD) differed between PDD and AD.
Vander Borght et al. (91)PDD 9
AD 9
C 9
[18F]fluorodeoxyglucose-PETIn PDD and AD hypometabolism with similar regional accentuation (lateral parietal, lateral temporal and lateral frontal association cortices and posterior cingulate cortex). In contrast to AD PDD showed greater metabolic reduction in the visual cortex and relatively preserved metabolism in the medial temporal cortex.
Gonzalez-Redondo et al. (92)PDND 14
PD-MCI 17
PDD 15
C 19
[18F]fluorodeoxyglucose-PETIn PD-MCI the hypometabolism exceeded atrophy in the angular gyrus, occipital, orbital and anterior frontal lobes. In PDD these areas were atrophic and surrounded by extensive hypometabolism.
Shinotoh et al. (93)PDND 14
PDD 2
PSP 12
C 13
Acetylcholinesterase activity using N-methyl-4-[11C]piperidyl acetate PETIn PDD higher reduction of choline acetyltransferase and acetylcholinesterase than in PDND.
Bohnen et al. (94)PDND 11
PDD 14
AD 12
C 10
Acetylcholinesterase activity using [11C]Methylpiperidin-4-ylpropionate PETMean cortical acetylcholinesterase activity was lowest in PDD.
Hiraoka et al. (95)PDD 12
C 13
[5-(11)C-methoxy]donepezil-PETIn PDD density of acetylcholinesterase in the cerebral cortices correlated with improvements in visuoperceptual function after 3 months of donepezil therapy.
Kotagal et al. (96)PDND 11
PDD 6
DLB 6
C 14
Acetylcholinesterase activity using [11C]Methylpiperidin-4-ylpropionate PETThalamic cholinergic denervation is present in PD, PDD, and DLB but not in AD.

Changes of function and connectivity related to cognitive impairment and dementia in Parkinson's disease.

PD, Patients with Parkinson's disease; PD-MCI, Parkinson's disease patients with mild cognitive impairment; PDD, Parkinson's disease patients with dementia; PDND, non-demented PD; DLB, Dementia with Lewy body; AD, Patients with Alzheimer's disease; C, Controls; MoCA, Montreal Cognitive Assessment; MMSE, Mini Mental Status Examination; PET, positron emission tomography.

For the assessment of white matter pathology using DTI and imaging of metabolites (Proton magnetic resonance spectroscopy) there is currently not enough longitudinal data available and the value of these techniques to predict cognitive decline has to be further explored. The existing studies indicate that microstructural changes, such as increased mean diffusivity or reduced fractional anisotropy in the hippocampus, the frontal and parietal white matter tracts are associated with cognitive decline in PD (68, 80, 81, 102104). In particular, an increased mean diffusivity may be predictive for cognitive decline before fractional anisotropy decreases. However, these findings need further validation in longitudinal studies.

Fatigue

Fatigue is a common symptom that includes both mental and physical aspects. Up to 70% of individuals with PD experience fatigue every day (105). Fatigue dramatically impairs quality of life (106). It is a complex syndrome emerging from dysfunction in the nervous, endocrine and immune system (107). From a clinical point of view fatigue is frequently associated with other non-motor syndromes, like sleepiness, apathy, depression and autonomic dysfunction (105, 108). However, fatigue can also occur as an isolated syndrome; it is therefore important to understand that fatigue and sleepiness or depression is not the same condition (109, 110). Central fatigue is commonly measured through questionnaires, such as the Fatigue Severity Scale (111) which is recommended by the Movement Disorder Society (MDS) task force (112). Central fatigue can be described as a feeling of constant exhaustion and can occur in various chronic disorders. Peripheral fatigue is characterized by failure to sustain the force of muscle contraction and is more readily accessible to quantification (106, 113).

A key mechanism underlying fatigue is the activation of the inflammatory cytokine network (107, 114). Therefore, inflammatory markers serve as potential biomarkers of fatigue. In particular, higher serum levels of IL-6, IL1-Ra, sIL-2R, and VCAM-1 were associated with higher fatigue levels in patients with newly diagnosed, drug-naïve PD (115, 116). This neuroinflammatory processes may promote glutamate dysregulation and further influence neuronal activity and neuroplasticity, and impact neuronal circuits mediating distress and motivation in PD (117119). Interestingly, higher serum uric acid levels were significantly associated with less fatigue (120).

In addition, dysfunction of the endocrine system, such as hypothalamic-pituitary-adrenal system which is connected to basal ganglia, amygdala, thalamus and frontal cortex, seems to contribute to the pathophysiology of fatigue (113). Although there are no neuropathological studies of PD-fatigue supporting this model so far, several neuroimaging studies showed that multiple brain areas are involved in fatigue in PD. These include frontal, temporal and parietal regions indicative of emotion, motivation and cognitive functions (121126). In SPECT imaging with technetium-99 hexamethyl-propylene-amine-oxime PD-fatigue was associated with reduced perfusion in the frontal lobe (125). Others used PET with dopaminergic and serotonergic markers in fatigued vs. non-fatigued PD patients. Less serotonergic marker binding was found in striatal and limbic regions (thalamus, anterior cingulate, amygdala, insula) in PD-fatigue. The striatal 18F-dopa uptake was similar in fatigued and non-fatigued groups, but voxel-based analysis localized the reduced dopamine uptake to the caudate and insula in PD-fatigue (127). In addition the serotonin transporter (SERT) availability was significantly reduced in the striatum and thalamus of fatigued PD patients, suggesting that increasing the brain level of serotonin may improve PD-fatigue (127). The reduced serotonergic transmission suggests that a disturbed neurotransmitter balance within the basal ganglia and associated regions changes the integration of emotional and motor information in limbic regions, thus resulting in fatigue symptoms (128). With regard to striatal dopamine transporter uptake, results are conflicting. Two studies found no difference between fatigued and non-fatigued PD (127, 129). In the study by Chou et al., striatal dopamine transporter uptake was a significant predictor of fatigue in mild but not moderate-to-severe PD. They postulated that the lack of association between fatigue and nigrostriatal loss in advanced PD may reflect a denervation “floor” effect (130). Many of these studies have assessed advanced disease stages and patients on dopaminergic treatment. In contrast, Tessitore et al. studied fatigue in drug-naïve early PD using resting-state functional MRI (fMRI). Fatigue itself, and fatigue severity were associated with a decreased connectivity within the supplementary motor area and an increased connectivity within the default mode network (121). Importantly, these functional abnormalities occur independently from both dopamine-induced connectivity and structural changes. This study is in line with earlier neurophysiological studies suggesting that abnormal premotor and primary motor cortices connectivity correlate with fatigue (131, 132). Tessitore et al. hypothesized that the increased connectivity of the default mode network represents an initial cognitive compensatory response to the fatigue-related motor connectivity changes. In this sense fatigued PD-patients, when internally oriented, have to increase mental expenditure to maintain the same level of motor planning performance in order to switch more easily to externally oriented processing (121).

In summary, abnormalities in motivation of self-initiated tasks and motor function may play a significant role in the pathophysiology of fatigue (133). While non-dopaminergic basal ganglia pathways seem to be involved in PD-fatigue, the dopaminergic dysfunction may only play a role through extrastriatal projections.

Depression

PD patients are twice as likely to develop depression compared to healthy individuals (134). Depressive symptoms affect 40–50% of PD patients and significantly impact quality of life in PD (2). In particular, patients with cognitive impairment, longer disease duration, motor fluctuations, female gender, and higher doses of levodopa are at risk to develop depression (9).

Like other NMS, depression seems to be linked to inflammatory signaling. Increased inflammatory responses have been described both in the brain and peripheral blood of PD patients (135). Depression correlated with a high serum level of IL-10 (136) and IL-6 (137). High levels of both sIL-2R and TNF-α in blood samples from PD patients were significantly associated with more severe depression and anxiety (119). As reflection of CNS involvement, high CRP levels in CSF of PD patients were associated with more severe symptoms of depression (32). However, these findings are not specific for PD. Chronic inflammation in physically ill patients is often associated with symptoms of depression and also occurs in normal aging (138140). Moreover, PD in general is characterized by elevated levels of inflammatory cytokines, such as IL-6, tumor necrosis factor, IL-1β, IL-2, IL-10, C-reactive protein, and RANTES (141).

Depression in PD is associated with several structural and functional changes in the limbic system. In particular, changes in the amygdala, hippocampus and orbitofrontal cortex were frequently reported in PD depression (142151). The involvement of the serotonergic system was demonstrated in post-mortem tissue and validated in vivo by several PET imaging studies (152155). Compared to controls the serotonin transporter binding in non-depressed PD was lower in the striatal region, the orbitofrontal cortex, and the dorsolateral pre-frontal cortex which is an area known to be involved in major depression (155). Using dopaminergic and serotonergic presynaptic transporter radioligands a prominent role of serotonergic degeneration in limbic regions such as the anterior cingulate cortex was demonstrated (156, 157). Other PET studies observed a higher availability of the serotonin transporter in the raphe nuclei and limbic regions of depressed PD patients (152, 153). Likewise, decreased plasma levels of serotonin were found to be correlated with severity of depression (158). However, studies of the serotonin metabolite 5-hydroxyindoleacetic acid (5-HIAA) in CSF from depressed and non-depressed PD patients, have yielded contradictory results (159), and serotonergic dysfunction alone may only explain vulnerability to depression in PD. Yet, symptoms of depression are also linked to mesolimbic dopaminergic degeneration (160, 161) which is in line with the clinical observation of improvement of depression by dopaminergic treatment (162).

Conclusion

From this overview emerges a comprehensive picture of recent fluid and imaging biomarkers which have been studied in a number of clearly defined and sizable cohorts of PD patients with PD. Especially longitudinal studies are necessary to make the biomarkers potentially useful for therapeutic or even clinical trial evaluation. A number of recent studies have provided ample evidence for specific predictive biomarkers across multiple domains combining clinical, biochemical, and neuroimaging information. Yet, at this stage a lack of standardized and comparable methods preclude clinical everyday use of these biomarkers beyond their value as diagnostic or prognostic tools in cohorts of patients. Thus, more research needs to be undertaken into finding reliable combinations of predictors of NMS in PD on an individual level, and standardization and harmonization of protocols in particular in CSF handling and neuroimaging has to be taken further.

Statements

Author contributions

TP and JG: conception, collection of data, interpretation of data, drafting the work; OW: revising the work critically for important intellectual content.

Acknowledgments

We thank Elena Huß for assistance of data collection.

Conflict of interest

The authors declare that the research was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest.

References

  • 1.

    SchapiraAHVChaudhuriKRJennerP. Non-motor features of Parkinson disease. Nat Rev Neurosci. (2017) 18:509. 10.1038/nrn.2017.91

  • 2.

    van UemJMMarinusJCanningCvan LummelRDodelRLiepelt-ScarfoneIet al. Health-related quality of life in patients with parkinson's disease–a systematic review based on the ICF model. Neurosci Biobehav Rev. (2016) 61:2634. 10.1016/j.neubiorev.2015.11.014

  • 3.

    Biomarkers Definitions Working Group. Biomarkers and surrogate endpoints: preferred definitions and conceptual framework. Clin Pharmacol Ther. (2001) 69:8995. 10.1067/mcp.2001.113989

  • 4.

    FenglerSLiepelt-ScarfoneIBrockmannKSchäfferEBergDKalbeE. Cognitive changes in prodromal Parkinson's disease: a review. Mov Disord. (2017) 32:165566. 10.1002/mds.27135

  • 5.

    AarslandDBrønnickKLarsenJPTysnesOBAlvesGNorwegian ParkWest Study Group. Cognitive impairment in incident, untreated Parkinson disease: the Norwegian ParkWest study. Neurology. (2009) 72:11216. 10.1212/01.wnl.0000338632.00552.cb

  • 6.

    LitvanIGoldmanJGTrösterAISchmandBAWeintraubDPetersenRCet al. Diagnostic criteria for mild cognitive impairment in Parkinson's disease: movement disorder society task force guidelines. Mov Disord. (2012) 27:34956. 10.1002/mds.24893

  • 7.

    AarslandDAndersenKLarsenJPLolkAKragh-SørensenP. Prevalence and characteristics of dementia in Parkinson disease: an 8-year prospective study. Arch Neurol. (2003) 60:38792. 10.1001/archneur.60.3.387

  • 8.

    SvenningssonPWestmanEBallardCAarslandD. Cognitive impairment in patients with Parkinson's disease: diagnosis, biomarkers, and treatment. Lancet Neurol. (2012) 11:697707. 10.1016/S1474-4422(12)70152-7

  • 9.

    MarinusJZhuKMarrasCAarslandDvan HiltenJJ. Risk factors for non-motor symptoms in Parkinson's disease. Lancet Neurol. (2018) 17:55968. 10.1016/S1474-4422(18)30127-3

  • 10.

    AnangJBGagnonJFBertrandJARomenetsSRLatreilleVPanissetMet al. Predictors of dementia in Parkinson disease: a prospective cohort study. Neurology. (2014) 83:125360. 10.1212/WNL.0000000000000842

  • 11.

    ZhuKvan HiltenJJMarinusJ. Predictors of dementia in Parkinson's dise findings from a 5-year prospective study using the SCOPA-COG. Parkinsonism Relat Disord. (2014) 20:9805. 10.1016/j.parkreldis.2014.06.006

  • 12.

    LitvanIAarslandDAdlerCHGoldmanJGKulisevskyJMollenhauerBet al. MDS Task Force on mild cognitive impairment in Parkinson's disease: critical review of PD-MCI. Mov Disord. (2011) 26:181424. 10.1002/mds.23823

  • 13.

    PaganoGDe MiccoRYousafTWilsonHChandraAPolitisM. REM behavior disorder predicts motor progression and cognitive decline in Parkinson disease. Neurology. (2018) 91:e894e905. 10.1212/WNL.0000000000006134

  • 14.

    Williams-GrayCHFoltynieTBrayneCERobbinsTWBarkerRA. Evolution of cognitive dysfunction in an incident Parkinson's disease cohort. Brain. (2007) 130(Pt 7):178798. 10.1093/brain/awm111

  • 15.

    KehagiaAABarkerRARobbinsTW. Cognitive impairment in Parkinson's disease: the dual syndrome hypothesis. Neurodegener Dis. (2013) 11:7992. 10.1159/000341998

  • 16.

    BraakHRübUJansen SteurENDel TrediciKde VosRA. Cognitive status correlates with neuropathologic stage in Parkinson disease. Neurology. (2005) 64:140410. 10.1212/01.WNL.0000158422.41380.82

  • 17.

    AarslandDPerryRBrownALarsenJPBallardC. Neuropathology of dementia in Parkinson's disease: a prospective, community-based study. Ann Neurol. (2005) 58:7736. 10.1002/ana.20635

  • 18.

    AlvesGLangeJBlennowKZetterbergHAndreassonUFørlandMGet al. CSF Aβ42 predicts early-onset dementia in Parkinson disease. Neurology. (2014) 82:178490. 10.1212/WNL.0000000000000425

  • 19.

    BäckströmDCErikssonDomellöf MLinderJOlssonBÖhrfeltATruppMet al. Cerebrospinal fluid patterns and the risk of future dementia in early, incident Parkinson disease. JAMA Neurol. (2015) 72:117582. 10.1001/jamaneurol.2015.1449

  • 20.

    BrockmannKLercheSDilgerSSStirnkorbJGApelAHauserAKet al. SNPs in Aβ clearance proteins: lower CSF Aβ1−42 levels and earlier onset of dementia in PD. Neurology. (2017) 89:233540. 10.1212/WNL.0000000000004705

  • 21.

    ComptaYMartíMJIbarretxe-BilbaoNJunquéCValldeoriolaFMu-ozEet al. Cerebrospinal tau, phospho-tau, and beta-amyloid and neuropsychological functions in Parkinson's disease. Mov Disord. (2009) 24:220310. 10.1002/mds.22594

  • 22.

    ComptaYEzquerraMMu-ozETolosaEValldeoriolaFRiosJet al. High cerebrospinal tau levels are associated with the rs57 tau gene variant and low cerebrospinal β-amyloid in Parkinson disease. Neurosci Lett. (2011) 487:16973. 10.1016/j.neulet.2010.10.015

  • 23.

    ComptaYPereiraJBRíosJIbarretxe-BilbaoNJunquéCBargallóNet al. Combined dementia-risk biomarkers in Parkinson's disease: a prospective longitudinal study. Parkinsonism Relat Disord. (2013) 19:71724. 10.1016/j.parkreldis.2013.03.009

  • 24.

    ComptaYValenteTSauraJSeguraBIranzoÁSerradellMet al. Correlates of cerebrospinal fluid levels of oligomeric- and total-α-synuclein in premotor, motor and dementia stages of Parkinson's disease. J Neurol. (2015) 262:294306. 10.1007/s00415-014-7560-z

  • 25.

    FfytcheDHPereiraJBBallardCChaudhuriKRWeintraubDAarslandD. Risk factors for early psychosis in PD: insights from the Parkinson's progression markers initiative. J Neurol Neurosurg Psychiatry. (2017) 88:32531. 10.1136/jnnp-2016-314832

  • 26.

    GmitterováKGawineckaJLlorensFVargesDValkovicPZerrI. Cerebrospinal fluid markers analysis in the differential diagnosis of dementia with Lewy bodies and Parkinson's disease dementia. Eur Arch Psychiatry Clin Neurosci. (2018). 10.1007/s00406-018-0928-9

  • 27.

    HalbgebauerSNaglMKlafkiHHaußmannUSteinackerPOecklPet al. Modified serpinA1 as risk marker for Parkinson's disease dementia: analysis of baseline data. Sci Rep. (2016) 6:26145. 10.1038/srep26145

  • 28.

    HallSÖhrfeltAConstantinescuRAndreassonUSurovaYBostromFet al. Accuracy of a panel of 5 cerebrospinal fluid biomarkers in the differential diagnosis of patients with dementia and/or parkinsonian disorders. Arch Neurol. (2012) 69:144552. 10.1001/archneurol.2012.1654

  • 29.

    HallSSurovaYÖhrfeltAZetterbergHLindqvistDHanssonO. CSF biomarkers and clinical progression of Parkinson disease. Neurology. (2015) 84:5763. 10.1212/WNL.0000000000001098

  • 30.

    HanssonOHallSOhrfeltAZetterbergHBlennowKMinthonLet al. Levels of cerebrospinal fluid α-synuclein oligomers are increased in Parkinson's disease with dementia and dementia with Lewy bodies compared to Alzheimer's disease. Alzheimers Res Ther. (2014) 6:25. 10.1186/alzrt255

  • 31.

    JanssensJVermeirenYFransenEAertsTVan DamDEngelborghsSet al. Cerebrospinal fluid and serum MHPG improve Alzheimer's disease versus dementia with Lewy bodies differential diagnosis. Alzheimers Dement. (2018) 10:17281. 10.1016/j.dadm.2018.01.002

  • 32.

    LindqvistDHallSSurovaYNielsenHMJanelidzeSBrundinLet al. Cerebrospinal fluid inflammatory markers in Parkinson's disease: associations with depression, fatigue, and cognitive impairment. Brain Behav Immun. (2013) 33:1839. 10.1016/j.bbi.2013.07.007

  • 33.

    MaetzlerWLiepeltIReimoldMReischlGSolbachCBeckerCet al. Cortical PIB binding in Lewy body disease is associated with Alzheimer-like characteristics. Neurobiol Dis. (2009) 34:10712. 10.1016/j.nbd.2008.12.008

  • 34.

    MaetzlerWStapfAKSchulteCHauserAKLercheSWursterIet al. Serum and cerebrospinal fluid uric acid levels in lewy body disorders: associations with disease occurrence and amyloid-β pathway. J Alzheimers Dis. (2011) 27:11926. 10.3233/JAD-2011-110587

  • 35.

    MaetzlerWTianYBaurSMGaugerTOdojBSchmidBet al. Serum and cerebrospinal fluid levels of transthyretin in Lewy body disorders with and without dementia. PLoS ONE. (2012) 7:e48042. 10.1371/journal.pone.0048042

  • 36.

    ModreanuRCerqueraSCMartíMJRíosJSánchez-GómezACámaraAet al. Cross-sectional and longitudinal associations of motor fluctuations and non-motor predominance with cerebrospinal τ and Aβ as well as dementia-risk in Parkinson's disease. J Neurol Sci. (2017) 373:2239. 10.1016/j.jns.2016.12.064

  • 37.

    ParnettiLTiraboschiPLanariAPeducciMPadiglioniCD'AmoreCet al. Cerebrospinal fluid biomarkers in Parkinson's disease with dementia and dementia with Lewy bodies. Biol Psychiatry. (2008) 64:8505. 10.1016/j.biopsych.2008.02.016

  • 38.

    ParnettiLFarottiLEusebiPChiasseriniDDe CarloCGiannandreaDet al. Differential role of CSF alpha-synuclein species, tau, and Aβ42 in Parkinson's Disease. Front Aging Neurosci. (2014) 6:53. 10.3389/fnagi.2014.00053

  • 39.

    SchragASiddiquiUFAnastasiouZWeintraubDSchottJM. Clinical variables and biomarkers in prediction of cognitive impairment in patients with newly diagnosed Parkinson's disease: a cohort study. Lancet Neurol. (2017) 16:6675. 10.1016/S1474-4422(16)30328-3

  • 40.

    SiderowfAXieSXHurtigHWeintraubDDudaJChen-PlotkinAet al. CSF amyloid β 1-42 predicts cognitive decline in Parkinson disease. Neurology. (2010) 75:105561. 10.1212/WNL.0b013e3181f39a78

  • 41.

    StewartTLiuCGinghinaCCainKCAuingerPCholertonBet al. Parkinson Study Group DATATOP Investigators. Cerebrospinal fluid α-synuclein predicts cognitive decline in Parkinson disease progression in the DATATOP cohort. Am J Pathol. (2014) 184:96675. 10.1016/j.ajpath.2013.12.007

  • 42.

    TerrelongeMJrMarderKSWeintraubDAlcalayRN. CSF β-amyloid 1-42 predicts progression to cognitive impairment in newly diagnosed parkinson disease. J Mol Neurosci. (2016) 58:8892. 10.1007/s12031-015-0647-x

  • 43.

    VranováHPHénykováEKaiserováMMenšíkováKVaštíkMMarešJet al. Tau protein, beta-amyloid1−42and clusterin CSF levels in the differential diagnosis of Parkinsonian syndrome with dementia. J Neurol Sci. (2014). 343:1204. 10.1016/j.jns.2014.05.052

  • 44.

    WennströmMSurovaYHallSNilssonCMinthonLBoströmFet al. Low CSF levels of both α-synuclein and the α-synuclein cleaving enzyme neurosin in patients with synucleinopathy. PLoS ONE. (2013) 8:e53250. 10.1371/journal.pone.0053250

  • 45.

    LiuCCholertonBShiMGinghinaCCainKCAuingerPet al. CSF tau and tau/Aβ42 predict cognitive decline in Parkinson's disease. Parkinson Relat Disord. (2015). 21:2716. 10.1016/j.parkreldis.2014.12.027

  • 46.

    MontineTJShiMQuinnJFPeskindERCraftSGinghinaCet al. CSF Aβ42 and tau in Parkinson's disease with cognitive impairment. Mov Disord. (2010) 25:26825. 10.1002/mds.23287

  • 47.

    PalmqvistSZetterbergHBlennowKVestbergSAndreassonUBrooksDJet al. Accuracy of brain amyloid detection in clinical practice using cerebrospinal fluid β-amyloid 42: a cross-validation study against amyloid positron emission tomography. JAMA Neurol. (2014) 71:12829. 10.1001/jamaneurol.2014.1358

  • 48.

    SkogsethREBronnickKPereiraJBMollenhauerBWeintraubDFladbyTet al. Associations between cerebrospinal fluid biomarkers and cognition in early untreated Parkinson's disease. J Parkinsons Dis. (2015) 5:78392. 10.3233/JPD-150682

  • 49.

    ZhouBWenMYuWFZhangCLJiaoL. The diagnostic and differential diagnosis utility of cerebrospinal fluid α-synuclein levels in Parkinson's disease: a meta-analysis. Parkinsons Dis. (2015) 2015:56738610.1155/2015/567386

  • 50.

    GaoLTangHNieKWangLZhaoJGanRet al. Cerebrospinal fluid alpha-synuclein as a biomarker for Parkinson's disease diagnosis: a systematic review and meta-analysis. Int J Neurosci. (2015) 125:64554. 10.3109/00207454.2014.961454

  • 51.

    StavALAarslandDJohansenKKHessenEAuningEFladbyT. Amyloid-β and α-synuclein cerebrospinal fluid biomarkers and cognition in early Parkinson's disease. Parkinsonism Relat Disord. (2015) 21:75864. 10.1016/j.parkreldis.2015.04.027

  • 52.

    BuddhalaCCampbellMCPerlmutterJSKotzbauerPT. Correlation between decreased CSF α-synuclein and Aβ1−42 in Parkinson diseaseNeurobiol Aging. (2015) 36:47684. 10.1016/j.neurobiolaging.2014.07.043

  • 53.

    HallSSurovaYÖhrfeltASwedish BioFINDER StudyBlennowKZetterbergHet al. Longitudinal measurements of cerebrospinal fluid biomarkers in Parkinson's Disease. Mov Disord. (2016) 31:898905. 10.1002/mds.26578

  • 54.

    LinCHYangSYHorngHEYangCCChiehJJChenHHet al. Plasma α-synuclein predicts cognitive decline in Parkinson's disease. J Neurol Neurosurg Psychiatry. (2017) 88:81824. 10.1136/jnnp-2016-314857

  • 55.

    Delgado-AlvaradoMGagoBNavalpotro-GomezIJiménez-UrbietaHRodriguez-OrozMC. Biomarkers for dementia and mild cognitive impairment in Parkinson's disease. Mov Disord. (2016) 31:86181. 10.1002/mds.26662

  • 56.

    AnnanmakiTPessala-DriverAHokkanenLMurrosK. Uric acid associates with cognition in Parkinson's disease. Parkinsonism Relat Disord. (2008) 14:5768. 10.1016/j.parkreldis.2007.11.001

  • 57.

    MocciaMPicilloMErroRVitaleCLongoKAmboniMet al. Is serum uric acid related to non-motor symptoms in de-novo Parkinson's disease patients?Parkinsonism Relat Disord. (2014) 20:7725. 10.1016/j.parkreldis.2014.03.016

  • 58.

    MocciaMPicilloMErroRVitaleCLongoKAmboniMet al. Presence and progression of non-motor symptoms in relation to uric acid in de novo Parkinson's disease. Eur J Neurol. (2015) 22:938. 10.1111/ene.12533

  • 59.

    PellecchiaMTSantangeloGPicilloMPivonelloRLongoKPivonelloCet al. Serum epidermal growth factor predicts cognitive functions in early, drug-naive Parkinson's disease patients. J Neurol. (2013) 260:43844. 10.1007/s00415-012-6648-6

  • 60.

    Chen-PlotkinASHuWTSiderowfAWeintraubDGoldmann GrossRHurtigHIet al. Plasma epidermal growth factor levels predict cognitive decline in Parkinson disease. Ann Neurol. (2011) 69:65563. 10.1002/ana.22271

  • 61.

    PellecchiaMTSantangeloGPicilloMPivonelloRLongoKPivonelloCet al. Insulin-like growth factor-1 predicts cognitive functions at 2-year follow-up in early, drug-naïve Parkinson's disease. Eur J Neurol. (2014) 21:8027. 10.1111/ene.12137

  • 62.

    FereshtehnejadSMZeighamiYDagherAPostumaRB. Clinical criteria for subtyping Parkinson's disease: biomarkers and longitudinal progression. Brain. (2017) 140:195976. 10.1093/brain/awx118

  • 63.

    PrellT. Structural and functional brain patterns of non-motor syndromes in Parkinson's Disease. Front Neurol. (2018) 9:138. 10.3389/fneur.2018.00138

  • 64.

    WeintraubDDietzNDudaJEWolkDADoshiJXieSXet al. Alzheimer's disease pattern of brain atrophy predicts cognitive decline in Parkinson's disease. Brain. (2012). 135(Pt 1):170180. 10.1093/brain/awr277

  • 65.

    MelzerTRWattsRMacAskillMRPitcherTLLivingstonLKeenanRJet al. Grey matter atrophy in cognitively impaired Parkinson's disease. J Neurol Neurosurg Psychiatry. (2012) 83:18894. 10.1136/jnnp-2011-300828

  • 66.

    LeeJEChoKHSongSKKimHJLeeHSSohnYHet al. Exploratory analysis of neuropsychological and neuroanatomical correlates of progressive mild cognitive impairment in Parkinson's disease. J Neurol Neurosurg Psychiatry. (2014) 85:716. 10.1136/jnnp-2013-305062

  • 67.

    BorroniBPremiEFormentiATurroneRAlbericiACottiniEet al. Structural and functional imaging study in dementia with Lewy bodies and Parkinson's disease dementia. Parkinsonism Relat Disord. (2015) 21:104955. 10.1016/j.parkreldis.2015.06.013

  • 68.

    DuncanGWFirbankMJYarnallAJKhooTKBrooksDJBarkerRAet al. Gray and white matter imaging: a biomarker for cognitive impairment in early Parkinson's disease?Mov Disord. (2016) 31:10310. 10.1002/mds.26312

  • 69.

    HattoriTOrimoSAokiSItoKAbeOAmanoAet al. Cognitive status correlates with white matter alteration in Parkinson's disease. Hum Brain Mapp. (2012) 33:72739. 10.1002/hbm.21245

  • 70.

    KandiahNZainalNHNarasimhaluKChanderRJNgAMakEet al. Hippocampal volume and white matter disease in the prediction of dementia in Parkinson's disease. Parkinsonism Relat Disord. (2014) 20:12038. 10.1016/j.parkreldis.2014.08.024

  • 71.

    RektorovaIBiundoRMarecekRWeisLAarslandDAntoniniA. Grey matter changes in cognitively impaired Parkinson's disease patients. PLoS ONE. (2014) 9:e85595. 10.1371/journal.pone.0085595

  • 72.

    BiundoRCalabreseMWeisLFacchiniSRicchieriGGalloPet al. Anatomical correlates of cognitive functions in early Parkinson's disease patients. PLoS ONE. (2013) 8:e64222. 10.1371/journal.pone.0064222

  • 73.

    PereiraJBSvenningssonPWeintraubDBrønnickKLebedevAWestmanEet al. Initial cognitive decline is associated with cortical thinning in early Parkinson disease. Neurology. (2014) 82:201725. 10.1212/WNL.0000000000000483

  • 74.

    HanganuABedettiCDegrootCMejia-ConstainBLafontaineALSolandVet al. Mild cognitive impairment is linked with faster rate of cortical thinning in patients with Parkinson's disease longitudinally. Brain. (2014) 137(Pt 4):11209. 10.1093/brain/awu036

  • 75.

    Ibarretxe-BilbaoNJunqueCSeguraBBaggioHCMartiMJValldeoriolaFet al. Progression of cortical thinning in early Parkinson's disease. Mov Disord. (2012) 27:174653. 10.1002/mds.25240

  • 76.

    MakESuLWilliamsGBFirbankMJLawsonRAYarnallAJet al. Baseline and longitudinal grey matter changes in newly diagnosed Parkinson's disease: ICICLE-PD study. Brain. (2015) 138(Pt 10):297486. 10.1093/brain/awv211

  • 77.

    HwangKSBeyerMKGreenAEChungCThompsonPMJanvinCet al. Mapping cortical atrophy in Parkinson's disease patients with dementia. J Parkinsons Dis. (2013) 3:6976. 10.3233/JPD-120151

  • 78.

    ZareiMIbarretxe-BilbaoNComptaYHoughMJunqueCBargalloNet al. Cortical thinning is associated with disease stages and dementia in Parkinson's disease. J Neurol Neurosurg Psychiatry. (2013) 84:87581. 10.1136/jnnp-2012-304126

  • 79.

    PagonabarragaJCorcuera-SolanoIVives-GilabertYLlebariaGGarcía-SánchezCPascual-SedanoBet al. Pattern of regional cortical thinning associated with cognitive deterioration in Parkinson's disease. PLoS ONE. (2013) 8:e54980. 10.1371/journal.pone.0054980

  • 80.

    CarlesimoGAPirasFAssognaFPontieriFECaltagironeCSpallettaG. Hippocampal abnormalities and memory deficits in Parkinson disease: a multimodal imaging study. Neurology. (2012) 78:193945. 10.1212/WNL.0b013e318259e1c5

  • 81.

    ChenBFanGGLiuHWangS. Changes in anatomical and functional connectivity of Parkinson's disease patients according to cognitive status. Eur J Radiol. (2015) 84:131824. 10.1016/j.ejrad.2015.04.014

  • 82.

    GorgesMMüllerHPLuléD; LANDSCAPE ConsortiumPinkhardtEHLudolphACet al. To rise and to fall: functional connectivity in cognitively normal and cognitively impaired patients with Parkinson's disease. Neurobiol Aging. (2015) 36:172735. 10.1016/j.neurobiolaging.2014.12.026

  • 83.

    BaggioHCSeguraBSala-LlonchRMartiMJValldeoriolaFComptaYet al. Cognitive impairment and resting-state network connectivity in Parkinson's disease. Hum Brain Mapp. (2015) 36:199212. 10.1002/hbm.22622

  • 84.

    AmboniMTessitoreAEspositoFSantangeloGPicilloMVitaleCet al. Resting-state functional connectivity associated with mild cognitive impairment in Parkinson's disease. J Neurol. (2015) 262:42534. 10.1007/s00415-014-7591-5

  • 85.

    TessitoreAEspositoFVitaleCSantangeloGAmboniMRussoAet al. Default-mode network connectivity in cognitively unimpaired patients with Parkinson disease. Neurology. (2012) 79:222632. 10.1212/WNL.0b013e31827689d6

  • 86.

    RektorovaIKrajcovicovaLMarecekRMiklM. Default mode network and extrastriate visual resting state network in patients with Parkinson's disease dementia. Neurodegener Dis. (2012) 10:2327. 10.1159/000334765

  • 87.

    Olde DubbelinkKTSchoonheimMMDeijenJBTwiskJWBarkhofFBerendseHW. Functional connectivity and cognitive decline over 3 years in Parkinson disease. Neurology. (2014) 83:204653. 10.1212/WNL.0000000000001020

  • 88.

    SeibertTMMurphyEAKaestnerEJBrewerJB. Interregional correlations in Parkinson disease and Parkinson-related dementia with resting functional MR imaging. Radiology. (2012) 263:22634. 10.1148/radiol.12111280

  • 89.

    LinWCChenPCHuangYCTsaiNWChenHLWangHCet al. Dopaminergic therapy modulates cortical perfusion in parkinson disease with and without dementia according to arterial spin labeled perfusion magnetic resonance imaging. Medicine. (2016) 95:e2206. 10.1097/MD.0000000000002206

  • 90.

    Le HeronCJWrightSLMelzerTRMyallDJMacAskillMRLivingstonLet al. Comparing cerebral perfusion in Alzheimer's disease and Parkinson's disease dementia: an ASL-MRI study. J Cereb Blood Flow Metab. (2014) 34:96470. 10.1038/jcbfm.2014.40

  • 91.

    Vander BorghtTMinoshimaSGiordaniBFosterNLFreyKABerentSet al. Cerebral metabolic differences in Parkinson's and Alzheimer's diseases matched for dementia severity. J Nucl Med. (1997) 38:797802.

  • 92.

    González-RedondoRGarcía-GarcíaDClaveroPGasca-SalasCGarcía-EulateRZubietaJLet al. Grey matter hypometabolism and atrophy in Parkinson's disease with cognitive impairment: a two-step process. Brain. (2014). 137(Pt 8):235667. 10.1093/brain/awu159

  • 93.

    ShinotohHNambaHYamaguchiMFukushiKNagatsukaSIyoMet al. Positron emission tomographic measurement of acetylcholinesterase activity reveals differential loss of ascending cholinergic systems in Parkinson's disease and progressive supranuclear palsy. Ann Neurol. (1999) 46:629. 10.1002/1531-8249(199907)46:1<62::AID-ANA10>3.0.CO;2-P

  • 94.

    BohnenNIKauferDIIvancoLSLoprestiBKoeppeRADavisJGet al. Cortical cholinergic function is more severely affected in parkinsonian dementia than in Alzheimer disease: an in vivo positron emission tomographic study. Arch Neurol. (2003) 60:17458. 10.1001/archneur.60.12.1745

  • 95.

    HiraokaKOkamuraNFunakiYHayashiATashiroMHisanagaKet al. Cholinergic deficit and response to donepezil therapy in Parkinson's disease with dementia. Eur Neurol. (2012) 68:137143. 10.1159/000338774

  • 96.

    KotagalVMüllerMLKauferDIKoeppeRABohnenNI. Thalamic cholinergic innervation is spared in Alzheimer disease compared to parkinsonian disorders. Neurosci Lett. (2012) 514:16972. 10.1016/j.neulet.2012.02.083

  • 97.

    Ramírez-RuizBMartíMJTolosaEBartrés-FazDSummerfieldCSalgado-PinedaPet al. Longitudinal evaluation of cerebral morphological changes in Parkinson's disease with and without dementia. J Neurol. (2005) 252:134552. 10.1007/s00415-005-0864-2

  • 98.

    MoralesDAVives-GilabertYGómez-AnsónBBengoetxeaELarra-agaPBielzaCet al. Predicting dementia development in Parkinson's disease using bayesian network classifiers. Psychiatry Res. (2013) 213:928. 10.1016/j.pscychresns.2012.06.001

  • 99.

    SchulzJPaganoGFernández BonfanteJAWilsonHPolitisM. Nucleus basalis of Meynert degeneration precedes and predicts cognitive impairment in Parkinson's disease. Brain. (2018) 141:150116. 10.1093/brain/awy072

  • 100.

    ArendtTBiglVArendtATennstedtA. Loss of neurons in the nucleus basalis of Meynert in Alzheimer's disease, paralysis agitans and Korsakoff's Disease. Acta Neuropathol. (1983) 61:1018. 10.1007/BF00697388

  • 101.

    CandyJMPerryRHPerryEKIrvingDBlessedGFairbairnAFet al. Pathological changes in the nucleus of meynert in Alzheimer's and Parkinson's diseases. J Neurol Sci. (1983) 59:27789. 10.1016/0022-510X(83)90045-X

  • 102.

    AgostaFCanuEStefanovaESarroLTomićAŠpicaVet al. Mild cognitive impairment in Parkinson's disease is associated with a distributed pattern of brain white matter damage. Hum Brain Mapp. (2014) 35:19219. 10.1002/hbm.22302

  • 103.

    TheilmannRJReedJDSongDDHuangMXLeeRRLitvanIet al. White-matter changes correlate with cognitive functioning in Parkinson's disease. Front Neurol. (2013) 4:37. 10.3389/fneur.2013.00037

  • 104.

    ZhengZShemmassianSWijekoonCKimWBookheimerSYPouratianN. DTI correlates of distinct cognitive impairments in Parkinson's disease. Hum Brain Mapp. (2014) 35:132533. 10.1002/hbm.22256

  • 105.

    FriedmanJHBrownRGComellaCGarberCEKruppLBLouJSet al. (2007). Working Group on Fatigue in Parkinson's Disease.Fatigue in Parkinson's disease: a review. Mov Disord. 22:297308. 10.1002/mds.21240

  • 106.

    FriedmanJHBeckJCChouKLClarkGFagundesCPGoetzCGet al. Fatigue in Parkinson's disease: report from a mutidisciplinary symposium. NPJ Parkinsons Dis. (2016) 2:15025. 10.1038/npjparkd.2015.25

  • 107.

    KlimasNGBroderickGFletcherMA. Biomarkers for chronic fatigue. Brain Behav Immun. (2012) 26:120210. 10.1016/j.bbi.2012.06.006

  • 108.

    ChouKLGilmanSBohnenNI. Association between autonomic dysfunction and fatigue in Parkinson disease. J Neurol Sci. (2017) 377:1902. 10.1016/j.jns.2017.04.023

  • 109.

    AlvesGWentzel-LarsenTLarsenJP. Is fatigue an independent and persistent symptom in patients with Parkinson disease?Neurology. (2004) 63:190811. 10.1212/01.WNL.0000144277.06917.CC

  • 110.

    van HiltenJJWeggemanMvan der VeldeEAKerkhofGAvan DijkJGRoosRA. Sleep, excessive daytime sleepiness and fatigue in Parkinson's disease. J Neural Transm Park Dis Dement Sect. (1993) 5:23544. 10.1007/BF02257678

  • 111.

    KruppLBLaRoccaNGMuir-NashJSteinbergAD. The fatigue severity scale. Application to patients with multiple sclerosis and systemic lupus erythematosus. Arch Neurol. (1989) 46:11213. 10.1001/archneur.1989.00520460115022

  • 112.

    KlugerBMHerlofsonKChouKLLouJSGoetzCGLangAEet al. Parkinson's disease-related fatigue: A case definition and recommendations for clinical research. Mov Disord. (2016) 31:62531. 10.1002/mds.26511

  • 113.

    ChaudhuriABehanPO. Fatigue in neurological disorders. Lancet. (2004) 363:97888. 10.1016/S0140-6736(04)15794-2

  • 114.

    BowerJE. Cancer-related fatigue: links with inflammation in cancer patients and survivors. Brain Behav Immun. (2007) 21:86371. 10.1016/j.bbi.2007.03.013

  • 115.

    HerlofsonKHeijnenCJLangeJAlvesGTysnesOBFriedmanJHet al. Inflammation and fatigue in early, untreated Parkinson's disease. Acta Neurol Scand. (2018) 138:3949. 10.1111/ane.12977

  • 116.

    PereiraJRSantosLVDSantosRMSCamposALFPimentaALde OliveiraMSet al. IL-6 serum levels are elevated in Parkinson's disease patients with fatigue compared to patients without fatigue. J Neurol Sci. (2016) 370:1536. 10.1016/j.jns.2016.09.030

  • 117.

    EyreHBauneBT. Neuroplastic changes in depression: a role for the immune system. Psychoneuroendocrinology. (2012) 37:1397416. 10.1016/j.psyneuen.2012.03.019

  • 118.

    MillerAHHaroonERaisonCLFelgerJC. Cytokine targets in the brain: impact on neurotransmitters and neurocircuits. Depress Anxiety. (2013) 30:297306. 10.1002/da.22084

  • 119.

    LindqvistDKaufmanEBrundinLHallSSurovaYHanssonO. Non-motor symptoms in patients with Parkinson's disease - correlations with inflammatory cytokines in serum. PLoS ONE. (2012) 7:e47387. 10.1371/journal.pone.0047387

  • 120.

    HuangXNgSYChiaNSAcharyyaSSetiawanFLuZHet al. Serum uric acid level and its association with motor subtypes and non-motor symptoms in early Parkinson's disease: PALS study. Parkinsonism Relat Disord. (2018) 55:5054. 10.1016/j.parkreldis.2018.05.010

  • 121.

    TessitoreAGiordanoADe MiccoRCaiazzoGRussoACirilloMet al. Functional connectivity underpinnings of fatigue in “Drug-Naïve” patients with Parkinson's disease. Mov Disord. (2016) 31:1497505. 10.1002/mds.26650

  • 122.

    ZhangJJDingJLiJYWangMYuanYSZhangLet al. Abnormal resting-state neural activity and connectivity of fatigue in Parkinson's disease. CNS Neurosci Ther. (2017) 23:2417. 10.1111/cns.12666

  • 123.

    LiJYuanYWangMZhangJZhangLJiangSet al. Alterations in regional homogeneity of resting-state brain activity in fatigue of Parkinson's disease. J Neural Transm. (2017) 124:118795. 10.1007/s00702-017-1748-1

  • 124.

    ChoSSAminianKLiCLangAEHouleSStrafellaAP. Fatigue in Parkinson's disease: The contribution of cerebral metabolic changes. Hum Brain Mapp. (2017) 38:28392. 10.1002/hbm.23360

  • 125.

    AbeKTakanashiMYanagiharaT. Fatigue in patients with Parkinson's disease. Behav Neurol. (2000) 12:1036. 10.1155/2000/580683

  • 126.

    ZhangLLiTYuanYTongQJiangSWangMet al. Brain metabolic correlates of fatigue in Parkinson's disease: a PET study. Int J Neurosci. (2018) 128:3306. 10.1080/00207454.2017.1381093

  • 127.

    PaveseNMettaVBoseSKChaudhuriKRBrooksDJ. Fatigue in Parkinson's disease is linked to striatal and limbic serotonergic dysfunction. Brain. (2010) 133:343443. 10.1093/brain/awq268

  • 128.

    PolitisMLoaneC. Serotonergic dysfunction in Parkinson's disease and its relevance to disability. Scientific World Journal. (2011) 11:172634. 10.1100/2011/172893

  • 129.

    SchifittoGFriedmanJHOakesDShulmanLComellaCLMarekKet al. Investigators. Fatigue in levodopa-naive subjects with Parkinson disease. Neurology. (2008) 71:4815. 10.1212/01.wnl.0000324862.29733.69

  • 130.

    ChouKLKotagalVBohnenNI. Neuroimaging and clinical predictors of fatigue in Parkinson disease. Parkinsonism Relat Disord. (2016) 23:459. 10.1016/j.parkreldis.2015.11.029

  • 131.

    LouJSBeniceTKearnsGSextonGNuttJ. Levodopa normalizes exercise related cortico-motoneuron excitability abnormalities in Parkinson's disease. Clin Neurophysiol. (2003) 114:9307. 10.1016/S1388-2457(03)00040-3

  • 132.

    BerardelliARothwellJCThompsonPDHallettM. Pathophysiology of bradykinesia in Parkinson's disease. Brain. (2001) 124(Pt 11):213146. 10.1093/brain/124.11.2131

  • 133.

    FabbriniGLatorreASuppaABloiseMFrontoniMBerardelliA. Fatigue in Parkinson's disease: motor or non-motor symptom?Parkinsonism Relat Disord. (2013) 19:14852. 10.1016/j.parkreldis.2012.10.009

  • 134.

    BeckerCBrobertGPJohanssonSJickSSMeierCR. Risk of incident depression in patients with Parkinson disease in the UK. Eur J Neurol. (2011) 18:44853. 10.1111/j.1468-1331.2010.03176.x

  • 135.

    Pessoa RochaNReisHJVanden BerghePCirilloC. Depression and cognitive impairment in Parkinson's disease: a role for inflammation and immunomodulation?Neuroimmunomodulation. (2014) 21:8894. 10.1159/000356531

  • 136.

    KarpenkoMNVasilishinaAAGromovaEAMuruzhevaZMBernadotteA. Interleukin-1β, interleukin-1 receptor antagonist, interleukin-6, interleukin-10, and tumor necrosis factor-α levels in CSF and serum in relation to the clinical diversity of Parkinson's disease. Cell Immunol. (2018) 327:7782. 10.1016/j.cellimm.2018.02.011

  • 137.

    VeselýBDufekMThonVBrozmanMKirálováSHalászováTet al. Interleukin 6 and complement serum level study in Parkinson's disease. J Neural Transm. (2018). 125:87581. 10.1007/s00702-018-1857-5

  • 138.

    BruunsgaardHPedersenMPedersenBK. Aging and proinflammatory cytokines. Curr Opin Hematol. (2001) 8:1316. 10.1097/00062752-200105000-00001

  • 139.

    ChenWWZhangXHuangWJ. Role of neuroinflammation in neurodegenerative diseases (Review). Mol Med Rep. (2016) 13:33916. 10.3892/mmr.2016.4948

  • 140.

    BritesDFernandesA. Neuroinflammation and depression: microglia activation, extracellular microvesicles and microRNA dysregulation. Front Cell Neurosci. (2015) 9:476. 10.3389/fncel.2015.00476

  • 141.

    QinXYZhangSPCaoCLohYPChengY. Aberrations in peripheral inflammatory cytokine levels in Parkinson disease: a systematic review and meta-analysis. JAMA Neurol. (2016) 73:131624. 10.1001/jamaneurol.2016.2742

  • 142.

    MatsuiHNishinakaKOdaMNiikawaHKomatsuKKuboriTet al. Depression in Parkinson's disease. Diffusion tensor imaging study. J Neurol. (2007) 254:11703. 10.1007/s00415-006-0236-6

  • 143.

    FeldmannAIllesZKosztolanyiPIllesEMikeAKoverFet al. Morphometric changes of gray matter in Parkinson's disease with depression: a voxel-based morphometry study. Mov Disord. (2008) 23:426. 10.1002/mds.21765

  • 144.

    KostićVSAgostaFPetrovićIGalantucciSSpicaVJecmenica-LukicMet al. Regional patterns of brain tissue loss associated with depression in Parkinson disease. Neurology. (2010) 75:85763. 10.1212/WNL.0b013e3181f11c1d

  • 145.

    SurdharIGeeMBouchardTCouplandNMalykhinNCamicioliR. Intact limbic-prefrontal connections and reduced amygdala volumes in Parkinson's disease with mild depressive symptoms. Parkinsonism Relat Disord. (2012) 18:80913. 10.1016/j.parkreldis.2012.03.008

  • 146.

    van MierloTJChungCFonckeEMBerendseHWvan den HeuvelOA. Depressive symptoms in Parkinson's disease are related to decreased hippocampus and amygdala volume. Mov Disord. (2015) 30:24552. 10.1002/mds.26112

  • 147.

    HuangCRavdinLDNirenbergMJPiboolnurakPSevertLManiscalcoJSet al. Neuroimaging markers of motor and nonmotor features of Parkinson's disease: an 18f fluorodeoxyglucose positron emission computed tomography study. Dement Geriatr Cogn Disord. (2013) 35:18396. 10.1159/000345987

  • 148.

    O'CallaghanCShineJMLewisSJHornbergerM. Neuropsychiatric symptoms in Parkinson's disease: fronto-striatal atrophy contributions. Parkinsonism Relat Disord. (2014) 20:86772. 10.1016/j.parkreldis.2014.04.027

  • 149.

    HuangPLouYXuanMGuQGuanXXuXet al. Cortical abnormalities in Parkinson's disease patients and relationship to depression: A surface-based morphometry study. Psychiatry Res Neuroimag. (2016) 250:248. 10.1016/j.pscychresns.2016.03.002

  • 150.

    CardosoEFMaiaFMFregniFMyczkowskiMLMeloLMSatoJRet al. Depression in Parkinson's disease: convergence from voxel-based morphometry and functional magnetic resonance imaging in the limbic thalamus. Neuroimage. (2009) 47:46772. 10.1016/j.neuroimage.2009.04.059

  • 151.

    LouYHuangPLiDCenZWangBGaoJet al. Altered brain network centrality in depressed Parkinson's disease patients. Mov Disord. (2015) 30:177784. 10.1002/mds.26321

  • 152.

    BoileauIWarshJJGuttmanMSaint-CyrJAMcCluskeyTRusjanPet al. Elevated serotonin transporter binding in depressed patients with Parkinson's disease: a preliminary PET study with [11C]DASB. Mov Disord. (2008) 23:177680. 10.1002/mds.22212

  • 153.

    PolitisMWuKLoaneCTurkheimerFEMolloySBrooksDJet al. Depressive symptoms in PD correlate with higher 5-HTT binding in raphe and limbic structures. Neurology. (2010) 75:19207. 10.1212/WNL.0b013e3181feb2ab

  • 154.

    BallangerBKlingerHEcheJLerondJValletAELe BarsDet al. Role of serotonergic 1A receptor dysfunction in depression associated with Parkinson's disease. Mov Disord. (2012) 27:849. 10.1002/mds.23895

  • 155.

    GuttmanMBoileauIWarshJSaint-CyrJAGinovartNMcCluskeyTet al. Brain serotonin transporter binding in non-depressed patients with Parkinson's disease. Eur J Neurol. (2007) 14:5238. 10.1111/j.1468-1331.2007.01727.x

  • 156.

    MailletAKrackPLhomméeEMétéreauEKlingerHFavreEet al. The prominent role of serotonergic degeneration in apathy, anxiety and depression in de novo Parkinson's disease. Brain. (2016) 139(Pt 9):2486502. 10.1093/brain/aww162

  • 157.

    SkidmoreFMYangMBaxterLvon DeneenKCollingwoodJHeGet al. Apathy, depression, and motor symptoms have distinct and separable resting activity patterns in idiopathic Parkinson disease. Neuroimage. (2013) 81:48495. 10.1016/j.neuroimage.2011.07.012

  • 158.

    TongQZhangLYuanYJiangSZhangRXuQet al. Reduced plasma serotonin and 5-hydroxyindoleacetic acid levels in Parkinson's disease are associated with nonmotor symptoms. Parkinsonism Relat Disord. (2015) 21:8827. 10.1016/j.parkreldis.2015.05.016

  • 159.

    SvenningssonPPålhagenSMathéAA. Neuropeptide Y and calcitonin gene-related peptide in cerebrospinal fluid in parkinson's disease with comorbid depression versus patients with major depressive disorder. Front Psychiatry. (2017) 8:102. 10.3389/fpsyt.2017.00102

  • 160.

    ThoboisSArdouinCLhomméeEKlingerHLagrangeCXieJet al. Non-motor dopamine withdrawal syndrome after surgery for Parkinson's disease: predictors and underlying mesolimbic denervation. Brain. (2010) 133(Pt 4):111127. 10.1093/brain/awq032

  • 161.

    KoertsJLeendersKLKoningMPortmanATvan BeilenM. Striatal dopaminergic activity (FDOPA-PET) associated with cognitive items of a depression scale (MADRS) in Parkinson's disease. Eur J Neurosci. (2007) 25:31326. 10.1111/j.1460-9568.2007.05580.x

  • 162.

    BaronePPoeweWAlbrechtSDebieuvreCMasseyDRascolOet al. Pramipexole for the treatment of depressive symptoms in patients with Parkinson's disease: a randomised, double-blind, placebo-controlled trial. Lancet Neurol. (2010) 9:57380. 10.1016/S1474-4422(10)70106-X

Summary

Keywords

Parkinson's disease, biomarker, non-motor syndromes, depression, fatigue, dementia

Citation

Prell T, Witte OW and Grosskreutz J (2019) Biomarkers for Dementia, Fatigue, and Depression in Parkinson's Disease. Front. Neurol. 10:195. doi: 10.3389/fneur.2019.00195

Received

01 October 2018

Accepted

15 February 2019

Published

08 March 2019

Volume

10 - 2019

Edited by

Tobias Warnecke, University Hospital Münster, Germany

Reviewed by

Walter Maetzler, University of Kiel, Germany; Gennaro Pagano, King's College London, United Kingdom

Updates

Copyright

*Correspondence: Tino Prell

This article was submitted to Movement Disorders, a section of the journal Frontiers in Neurology

Disclaimer

All claims expressed in this article are solely those of the authors and do not necessarily represent those of their affiliated organizations, or those of the publisher, the editors and the reviewers. Any product that may be evaluated in this article or claim that may be made by its manufacturer is not guaranteed or endorsed by the publisher.

Outline

Cite article

Copy to clipboard


Export citation file


Share article

Article metrics