Abstract
At the beginning of the 20th century, Knudsen illustrated that the mean observed salinity of the Baltic Sea could be realistically estimated, assuming an inflow of saline Kattegat water equals the net freshwater supply, also called the Knudsen theorem. As given in the historical review, several studies have followed the approach of well-mixed boxes, including time variations and a division between different sub-basins in the Baltic Sea. The box concept was later developed into mechanistic models by resolving the vertical structure in each sub-basin and adding processes related to vertical mixing, strait flow dynamics, and exchange with the atmosphere. However, as with the box concept, each sub-basin was assumed to be horizontally homogeneous. Early on, it was clear that the Baltic Sea circulation was highly unsteady, with fronts and eddies at different scales, illustrating a typical marine turbulent flow with energy cascade from basin scale to mesoscale, submesoscale, and microscale, where the energy dissipates. Many observational and modeling studies addressing the three-dimensional structure were developed over the last half-century. The approach of mechanistic models is useful for interpreting large-scale effects of meso- and submesoscale processes and for climate and long-term studies. The submesoscale approaches, including in situ observations, remote sensing, and models resolving the three-dimensional structure, may guide parametrizations of exchange between and within the different sub-basins. Recent submesoscale studies suggest localized eddy-rich regions: Arkona Basin, Gulf of Finland, Irbe Strait, Åland Sea connections, and several coastal areas.
1 Introduction
The Baltic Sea does not have strong currents like the Gulf Stream or Kuroshio that guide essential parts of the ocean water cycle. Instead, the water balance with a surplus of freshwater flow, combined with estuarine salinity gradients, forms mean water exchange flows in the straits connecting the sea with the world ocean or in the straits and deep channels connecting the morphometric subdivisions of the sea. At the same time, sea basins are stratified but horizontally relatively homogeneous, meaning that hydrographic contrasts within the basins are generally smaller than temperature and salinity differences between the basins. Among the water budget components, the inflow of saline water from the North Sea is highly variable. Already in early studies, Kalle (1943) found that a large inflow pulse in 1933/1934 caused an abrupt salinity increase in deep basins like the Eastern Gotland Basin (Figure 1), as noted by .
FIGURE 1
Knowledge of governing Baltic Sea water exchange processes has evolved in concert with developments in ocean studies. It is essential for understanding processes in the global and regional Earth System dynamics; using knowledge about the physical system forms the basis for climate and environmental studies. Rather long ago,
Specific to the Baltic Sea, hydrographic studies have been coordinated since the 1900s by the International Council for the Exploration of the Sea (ICES) to support fisheries regulations (Leppäranta and Myrberg, 2009). In the 1970s and 1980s, studies that specialized in monitoring the deep basins (
The Baltic Sea water cycle was extensively studied in the 20th century, and major research efforts were made in the
In the late 1980s, the Global Energy and Water Experiment (GEWEX) was developed within the World Climate Research Programme (WCRP) framework. The aims were to better understand global, regional, and local processes that exchange energy and water in the climate system. Here, the Baltic Sea served as one of the major sea regions for developing new measuring and modeling efforts. The box model concept served as a conceptual view of the climate problem (Figure 2), and several studies were developed, including observations and models of different complexities. These studies were later analyzed in several reviews (Omstedt et al., 2004; Reckermann et al., 2011; Omstedt et al., 2014; Omstedt and von Storch, 2023). Within the Baltic Earth Assessment Reports (Meier et al., 2023), recent knowledge of Baltic Sea salinity dynamics and related water exchange issues has been overviewed by Lehmann et al. (2022).
FIGURE 2

BALTEX I box presents a conceptual view of the Baltic Sea region’s coupled atmosphere, ocean, and land system. The arrows indicate different types of interactions between the boxes (figure courtesy of Marcus Reckermann).
Since the 1980s, different research schools of physical oceanography - the budget approach and meso-to-small scale approach – emerged from observation and modeling technology developments. Initially, these approaches were rather separate, but they have continued to converge in recent years. The Baltic Earth program has stimulated progress in such a convergence. The present review aims at a joint analysis and outline of historical developments of the budget and meso-to-small scale approaches, including submesoscale. In particular, the evolution of research hypotheses and societal needs, combined with enabling technologies, is presented. We also discuss gaps in the knowledge and propose some contemporary research questions.
2 Marine science organization background: need for budgets of water and chemicals
The interest in exploring the sea increased in the 19th century with inspiration from the Challenger expedition (1872–1876), under the command of Charles Wyville Thomson and supported by the British Royal Navy. Expeditions in coastal seas, such as the Baltic Sea, started. In the summer of 1877, Fredrik Laurentz Ekman made temperature and salinity measurements around Sweden, from the Skagerrak to the Bothnian Bay (
Water exchange heavily impacts the fate of persistent pollutants like heavy metals, Polychlorinated biphenyls (PCBs), and dioxins, which are deposited into the sea. Pollution of the seas became on the agenda in the 1960s. Jensen et al. (1969) demonstrated that in the Baltic Sea, the levels of chlorinated hydrocarbons are approximately ten times greater than for comparable species in the North Sea area and the Atlantic. In 1969, ICES formed the Working Group on Pollution of the Baltic, which, in the report published in 1974, outlined that “… basic hydrodynamical studies of the mechanisms for exchange and transfer of matter in the Baltic are of prime importance … ” (
Pollution by excess load of nutrients causes eutrophication, resulting in massive algal blooms and oxygen deficiency, controlled mainly by the vertical exchange of nutrients and temporal weakening of lateral deep-water transport of oxygen-rich North Sea waters (Jansson, 1978). The initial goal was to limit the phosphorus and nitrogen loads by 50% in all the sea areas (
The research landscape of the Baltic Sea has many aspects, as presented by
3 Mechanistic models of connected sub-basins
Mechanistic models (in their simplest forms, also called box models) start from water and salt conservation laws, including prescribed values on in- and outflows. Early in the 20th century, Knudsen (1900) illustrated that the mean observed salinity of the Baltic Sea could be realistically estimated, assuming an inflow of saline Kattegat water equals the net freshwater supply, also called the Knudsen theorem. Several model studies of water transport have followed, including time variations and a division between the Öresund and the Great Belt (e.g.,
Based on the stationary Knudsen theorem, water budgets were developed for the different sub-basins of the Baltic Sea (SNV, 1988;
FIGURE 3

A scheme of stationary water exchange in the Baltic Sea, including the sub-basins Baltic Proper, Bothnian Sea, and Bothnian Bay. The flows in the Baltic Proper also include the Gulf of Finland and the Gulf of Riga. From: The marine environment of Sweden - ecosystems under pressure. Flows in km3 per year. Adopted from SNV (1988).
The box concept was later developed into mechanistic models by adding more basic scientific laws related to vertical mixing, strait flow dynamics, and exchange with the atmosphere. As the first step, Stigebrandt (1983) developed a mechanistic model for the Baltic Sea entrance area. The dynamic in the Baltic Sea entrance area was modeled by only considering the Kattegat and the Belt Sea, both modeled as horizontally homogeneous two-layer sub-models. The model was driven by fresh water supply to the Baltic Sea and sea level variations between the Kattegat and the Baltic Sea, forcing the barotropic and baroclinic exchange flows. The model captured the main features of temporal salinity variations, indicating the validity of applied water exchange formulations. This model was connected with one strait only; studies by Omstedt (1987),
To include stratification effects within the Baltic Sea sub-basins, Omstedt et al. (1983) and Stigebrandt (1985) developed vertical one-dimensional pycnocline sub-models for temperature and salinity, driven by surface boundary conditions and lateral transports from the neighboring sub-basins. In the stratified water column below the vertically mixed layer, the dense flows from the upstream basin sink to the level of neutral buoyancy and move the overlying layers upward (Stigebrandt, 1985). The mixed layer model covers both the downward entrainment and retreat of the pycnocline. The importance of considering both the temperature and salinity stratification was demonstrated by a study of the cooling of surface water in the Bothnian Bay (Omstedt et al., 1983). Such a model was later used for spin-up simulations, showing that current Baltic Sea conditions could be realistically modeled after the spin-up period, starting from arbitrary initial conditions (Omstedt and Hansson, 2006). Namely, under the forcing by freshwater input and calculated water exchange between the sub-basins, the numerical experiment on the reconstruction of the present-day temperature and salinity regime, starting from the oceanic salinity in the whole Baltic Sea in 1700, reached the contemporary salinities in about 100 years (Figure 4).
FIGURE 4

Calculated surface salinity spin-up (grey) from ocean initial conditions using the PROBE-Baltic model of connected basins and observation (green). The forcing data are based on a 30-year period, which was repeated 10 times to reconstruct the 300-year time series. Adopted from Omstedt and Hansson (2006).
Mechanistic models later address the connection of different sub-basins in the Baltic Sea (e.g., Omstedt, 1990; Savchuk et al., 2012), the deep water ventilation in and between the different sub-basins (e.g., Stigebrant, 1987; Kõuts an Omstedt, 1993; Marmefelt, and Omstedt, 1993), sea ice (e.g.,
The mechanistic models assume that changes in salinity and other model variables along the estuarine gradients are much larger between the basins than within the basins, allowing horizontal integration of variables over these sea basins. Observational evidence for such an assumption stems partly from the long-term monitoring data (e.g., Kõuts and Omstedt, 1993). In the mechanistic models, the exchange flows of water and substances (including salt and heat) between the basins are parameterized by the density and sea level differences. Atmospheric data are also used when appropriate. Generally, the barotropic exchange is calculated using water-level forcing from the Kattegat and river runoff. In straits wider than the local internal Rossby radius, the baroclinic outflows are assumed to be in geostrophic balance (Stigebrandt, 1983; Omstedt, 1990). In narrow straits, such as between the Bothnian Bay and the Bothnian Sea, the baroclinic exchange is considered to be at a maximum flow rate calculated from baroclinic hydraulic control (Omstedt and Axell, 2003).
High-resolution observations, whose data are available from the 1980s, were used to check the horizontally integrated model’s assumptions of a small ratio of within-basin to between-basin salinity variations. Shipborne quasi-synoptic aerial surveys (∼1 day, ∼50 km) on eddy-resolving grids in two regions – the Eastern Gotland Basin and the Bornholm Basin, revealed that large-scale salinity variance between the basins (∼100 km) highly dominates over the mesoscale (∼10 km) spatial variance within one mapping, and the temporal (∼ few days) variability from one mapping to another (Kahru and Aitsam, 1985). Long sections of FerryBox data, measured onboard regularly cruising ships with a resolution of less than 1 km since the beginning of the 1990s (Karlson et al., 2016), have also revealed that surface salinity variations within the basins are much smaller than salinity drops in the frontal areas between the basins, especially between the Bothnian Sea and the Bothnian Bay. Regarding the depth distribution of variability, basin-to-basin salinity differences are larger in the deep layers below the halocline (depth >60 m) than in the layers above.
The classical Knudsen theorem for stationary flows has recently been revisited with high-resolution time-dependent data by
4 Basin-scale dynamics, fronts and upwelling
The Baltic Sea circulation is highly unsteady, as revealed already from historical observations. Time series of currents are often dominated by 14-h inertial oscillations, first recorded by
Deep-water flows below the halocline, located in the Baltic Proper at about 60 m depth, generally follow the right-hand slopes of deep sub-basins on its way from the Danish Straits (from Kattegat to Belt Sea and Öresund) to the Western Gotland Basin (Figure 1). Deep layers undergo large variations in flow, salinity and oxygen due to sporadically occurring large inflow pulses of saline water from the North Sea (Matthäus and Franck, 1992; Mohrholz, 2018), termed Major Baltic Inflow (MBI). Figure 5 gives an example of deep water spreading from the Bornholm Basin through the Stolpe Channel to the Eastern Gotland Basin after the November 2002 MBI, using numerical experiments with a tracer (Meier, 2007). There is complicated meso-to-small scale variability of deep water exchange in constrictions like the Bornholm Strait (Petrén and Walin, 1976;
FIGURE 5

Monthly mean tracer concentration after the Major Baltic Inflow of the North Sea water in November 2002, evolved to January 2003 (A) and March 2003 (B). The results were taken from an experiment when the tracer was initialized in the deep layers of the Bornholm Basin. Adopted from Meier (2007).
Fronts as high-gradient zones are formed when flows between the larger water masses converge, enhancing the horizontal gradients due to frontogenesis (Simpson, 1981; McWilliams, 2021). Such converging flows are often in geostrophic balance according to the Margules’ formula. The width of the frontal current jet is usually smaller than the Rossby deformation radius, but its length may extend to the dimensions of the basins. The frontal zones perform excursions due to meandering and wind drift. The variations within the estuarine salinity gradient are the strongest in the Danish Straits, where salinity drops from about 30 to 8–10 psu (Voss et al., 2011; Lehmann et al., 2022). (We adopted the practical salinity unit psu from historical data and/or figures; for reference, 10 psu corresponds to the absolute salinity of about 10.87 g kg-1). To the east of these straits, constrictions in the sea topography guide circulation and mixing (Figure 1) that affect the mean surface salinity distribution (Figure 6A). Outside the North Sea-Baltic Sea transition area, permanent forth-and-back migrating fronts are the Quark Front between the Bothnian Sea and Bothnian Bay (salinity drop between the areas from 1 to 3 psu,
FIGURE 6

Long-term annual mean salinity (A) and its temporal standard deviation (B) for the period 1993–2019, based on the CMEMS daily reanalysis data (www.copernicus.eu/en/access-data/copernicus-services-catalogue/baltic-sea-physics-reanalysis). Color scales were adjusted for the areas east of the Arkona Basin. Instead of historical salinity unit psu, the absolute salinity g kg-1 is used as in the model data. From Suursaar et al. (2021).
Fronts also appear in the temperature fields (Kahru et al., 1995;
Upwelling occurs when wind-induced Ekman transport drifts surface waters away from the coast, and deeper waters reach the surface (Lehmann and Myrberg, 2008). Such wind situations are frequently found on the Swedish south and east coasts, the Swedish coast of Bothnian Bay, the southern tip of Gotland, and the Finnish coast of the Gulf of Finland (Lehmann et al., 2012). Upwelling also occurs off the Estonian coast and the Baltic east coast, the Polish coast, and the west coast of Rügen. In the Gulf of Finland, the upwelling waters may cover up to 38% of the gulf’s surface area, while the filaments may cover up to 5% (Uiboupin and Laanemets, 2009). The duration of the upwelling depends on the duration of favorable winds; relaxation after the winds become unfavorable may take several days. Recent knowledge of upwelling in the Baltic Sea has been summarized by Lehmann et al. (2022).
A 2-week upwelling event was observed in August 2006 along the Estonian coast during easterly to southeasterly winds. It was extensively studied using in situ observations (Lips et al., 2009) and modeling combined with remote sensing (Laanemets et al., 2011). Consecutive sea source temperature maps (Figure 7) obtained from satellite imagery and numerical modeling revealed that the offshore drift of the outcropped thermocline was modulated by meandering mesoscale coherent structures (current squirts, filaments). The general features of the observed patterns were modeled well. Individual mesoscale features had uncertainties in modeling the timing and location due to the random nature of eddy and filament generation. Over 2 months, including the upwelling period, mesoscale currents comprised 66% of the total kinetic energy (corresponding to r.m.s. (root-mean-square) velocity fluctuations from 0.14 to 0.20 m s-1), while inertial oscillations with a 14-h period contained 20%. By comparing the amounts of upper-layer nutrients, it was found that during the upwelling-dominated period, the excess vertical transport of phosphorus by upwelling was comparable to the external land-based load.
FIGURE 7

Sea surface temperature maps of the Gulf of Finland in August 2006: satellite imagery (A–D), model (E–H). Adopted from Laanemets et al. (2011).
5 Mesoscale dynamics
Mesoscale eddies are swirling patterns of currents of nearly circular shape capable of traveling as compact features over distances larger than their size. The Rossby deformation radius Rd is a key to defining the horizontal scale of mesoscale motions. It is the product of mean stratification strength (Väisälä-Brunt frequency) and depth divided by the Coriolis parameter, reflecting the effect of Earth rotation at a given latitude. Mesoscale eddies typically have a diameter a few times the Rd in the ocean and the marginal seas (
Mesoscale eddies comprise an important intermediate part of the multitude of physical features in the sea, generated by the cascade of interacting physical processes, from the basin scales to the turbulent mixing scales. Within the basins, mesoscale eddies are formed by instability processes or forced baroclinic flows crossing the depth contours. Mesoscale eddies, which have a dominating geostrophic balance between the currents and horizontal pressure gradients (Rossby number Ro < 1), decay into ageostrophic (Ro > 1) submesoscale eddies that cause isopycnal and diapycnal mixing, with the help of internal waves and the thermohaline fine structure. Mesoscale eddies cause a “streaky” distribution of sea surface temperature and plankton variables (
FIGURE 8

Cyanobacterial (primarily Nodularia spumigena) accumulations in the Northern Baltic Proper on 11 July 2005 as shown on MODIS Terra quasi true color image at 250 m resolution. Adopted from Kahru and Elmgren (2014).
Oceanic mesoscale eddies were discovered in the 1970s as persistent, slowly evolving eddy structures in current distributions (Koshlyakov and Grachev, 1973; McWilliams, 1976). The eddy currents are in geostrophic balance with density variations. Historical hydrographic data processing has often classified large temperature, salinity, and density anomalies as “suspect”. After discovering the mesoscale eddies, the abandoned density anomalies were used to construct worldwide maps of eddy occurrence (
Remote sensing is a valuable tool for identifying eddies over larger regions and studying their properties, as noted at the beginning of eddy science (Robinson, 1983). For the Baltic Sea,
Altimetry has a good dynamic presentation since it determines sea level gradients, enabling geostrophic calculations of ocean currents. The present altimetry is good for resolving oceanic eddies with diameters 100–200 km and a lifetime of more than 4 weeks (
Regarding the historical view on mesoscale variability, phytoplankton patchiness was found in the Baltic Sea in the 1970s. It stimulated the planning of an interdisciplinary study of mesoscale dynamics (ICES, 1979). The multi-ship (13 research vessels) 2-week patchiness study experiment PEX-86 was conducted during the spring bloom period (April - May) in the southern part of the Eastern Gotland Basin on a study grid of 20 x 40 nautical miles (
FIGURE 9

Observational data from 7 May 1986. (A) Map of the dynamic height of 10 dbar relative to 90 dbar, in dynamic cm (solid lines) and depth contours (dashed lines) around the map area. The position of the lower panel transect (B) is given by a heavy broken line. (B) A 40-mile salinity transect in the depth range 0–100 m, grid step between the profiles 4 nautical miles. The contour interval is 0.2 psu for solid lines and 0.02 psu for dashed lines. Modified from
Advanced in situ observations of eddies (Zhurbas and Paka, 1999; Pavelson et al., 1999; Stigebrandt et al., 2002; Piechura and Beszczynska-Möller, 2004; Lass and Mohrholz, 2003; Lass et al., 2003; Voss et al., 2005;
The properties of mesoscale eddies throughout their life cycle were studied by applying eddy detection algorithms to data from high-resolution numerical modeling. Vortmeyer-Kley et al. (2019) used three different eddy detection methods based on Eulerian and Lagrangian approaches to analyze the modeled current fields with a 0.6 km resolution. Over the 2-year modeling period, about 100,000 eddies were detected. In the variety of eddies, the lifetime of more than 2 days is covered by about 10% of eddies, but short-living eddy structures with lifetimes less than the inertial period (about 14 h) comprise about 40% of the eddy detections (however, it depends on the method used). On average, the eddies detected in the model have a diameter of 16 km, whereas the maximum diameter amounted to 40 km. Most eddies travel about 20 km, but migrations up to 80 km were also found. An independent example of long-time eddy travel for 33 days is given by Väli et al. (2017). Travkin et al. (2024) used the Baltic Sea Physics Reanalysis 1993–2020 with a 2 km resolution. Compared to the Rossby deformation radius, such a large grid step is known to smear out smaller eddies. However, similar to Vortmeyer-Kley et al. (2019), they found that most eddies have a diameter of 10–20 km, a lifetime of 2–3 days, and a sea level anomaly of 0.05–0.20 m in the eddy center.
Both the eddy detection studies indicated the dominance of cyclones at the surface and increased eddy activity during late autumn and winter, as was earlier found from in situ observations (Kõuts et al., 1990) and satellite SAR images (Karimova, 2012). Statistical detections of eddy diameters, lifetimes, and travel distances follow the results from observations. Persistent eddies observed at the surface are related to 10–20 m vertical excursions of halocline due to the geostrophic relations at low Rossby numbers. Eddies related to the isopycnal displacements in the thermocline have shorter lifetimes and smaller diameters.
Subsurface eddies have a belt of maximum vortex currents either in the halocline, the thermocline, or the intermediate layer between these layers. Intra-halocline lenslike eddies, identified from observations (
The eddy structures, found either in observational data or numerical results, need to be interpreted in terms of theoretical solutions, establishing the dependence between the parameters. In the ocean, the first observations of mesoscale eddies were interpreted as slowly moving linear Rossby waves (e.g., Koshlyakov and Grachev, 1973;
The nature of Baltic Sea mesoscale eddies can be divided by generation mechanisms - forced or random eddies. Forced eddies occur due to wind action when a variable larger-scale flow crosses the (abruptly changing) depth contours or goes past the coastal capes. Their size is determined mainly by topographic features and stratification. Under similar weather forcing, the eddies are formed in nearly the same locations. Random eddies are generated due to the baroclinic and barotropic instability of disturbances that may grow in a sheared flow. Their size is a few times the Rossby deformation radius. During their life cycle, from formation to decay, eddies may migrate as compact features over significant distances.
Eddies comprise a significant chain in the energy and vorticity cascade of multi-scale physical transport and mixing processes (Meier et al., 2006; Reissmann et al., 2009), enhancing them at large spatial and temporal scales. Mesoscale eddies cause relatively high values of lateral diffusion coefficients, from 500 to 2000 m2 s-1 (Zhurbas et al., 2008;
6 Submesoscale dynamics and mixing
Submesoscale dynamics cover motions with a scale smaller than Rossby’s deformation radius (mesoscale) and larger than the thermohaline fine structure and turbulent mixing. Oceanic submesoscale patterns have a scale of 200 m–20 km (Taylor and Thompson, 2023). Submesoscale variability is seen on the sea surface as squirts and striped structures of temperature or other scalar fields, often curved along mesoscale eddies and fronts. While geostrophic balance dominates the mesoscale motions, ageostrophic effects are important in submesoscale dynamics. Oceanic submesoscale features significantly affect large-scale flows’ transport, mixing, and dissipation (McWilliams, 2019). Knowledge of submesoscale dynamics has advanced during the recent decade thanks to the rapid developments in observation and modeling techniques. Earlier knowledge on thermohaline intrusions (Ruddick and Richards, 2003) has been merged into the submesoscale concept.
In the Baltic Sea, the submesoscale processes with high values of Rossby number have spatial scales across the intrusive thread of less than 5 km and time scales of a few days (Lips et al., 2016; Väli et al., 2017;
FIGURE 10

(A–C) Buoyancy along the glider transect at 3, 15, and 25 m depth on 20–23 May 2018. (D–F) Temperature distributions (black contours with a step of 1°C) as a function of depth (0–25 m) overlaid with white contours marking the relative geostrophic velocity with a step of 2.5 cm s-1 on 20–23 May 2018. The sampling timeline is at the top of the columns. The left side of a subplot is located closer to the coast and presents the southern part of the section. The distance between the ticks presenting latitude is about 2.7 km. (G–I) Temperature deviations as a function of pressure along the glider section on 20–23 May 2018. Black contours mark potential density anomaly with a step 0.1 kg m-3. (J–L) Isopycnal temperature distributions (black contours with a step of 1°C) along the glider section on 20–23 May 2018. From Salm et al. (2023).
Numerical models, resolving the submesoscale with a grid step of 0.6 km or less, enable comprehensive dynamical analysis of the terms and balances in the hydrodynamic equations. A useful approach to handle the submesoscale structures is to calculate “gradient” characteristics. The Rossby number (Ro), as a ratio of the vertical vorticity component to the Coriolis parameter, is one of the main objects of such analysis, distinguishing between the geostrophic mesoscale (Ro < 1) and ageostrophic submesoscale (Ro > 1) regimes. Other quantities to explain the fate of threads include the horizontal divergence of currents, the horizontal buoyancy gradient, vertical velocity, Richardson number, frontogenetic strain rate, thermohaline spice (defined as the ratio of temperature and salinity variations to yield unchanged water density), but also kinetic energy and potential energy anomaly (Väli et al., 2017; Onken et al., 2020;
A study by
FIGURE 11

Maps of the surface dynamics in the Eastern Gotland Basin (between the Gotland island on the west and the Latvian coast on the east) on 19 October 2017. (A) Sea surface temperature (SST), (B) Rossby number, i.e., relative vorticity normalized by the Coriolis frequency. Modified from
Model studies of the eddy statistics over annual or longer periods (Vortmeyer-Kley et al., 2019; Väli et al., 2024) have revealed the dominance of cyclonically curved submesoscales over anticyclonic. Preference for cyclonic vorticity has also been pointed out by shorter model studies (Väli et al., 2017; Onken et al., 2020; Zhurbas et al., 2022) and SAR satellite images (Karimova, 2012). It was also found that there was a higher intensity of mesoscale eddies and submesoscale features during the winter compared to the summer. Väli et al. (2024) concluded that the vertically averaged kinetic energy (including that of meso- and submesoscales) is about 70 cm2 s-2 in December–January and is reduced to 30 cm2 s-2 in June–July. For the observational background, deep-layer available potential energy was calculated from the mesoscale CTD surveys conducted in 1984–1992 in the Eastern Gotland Basin. Analysis of vertical excursions of isopycnals and the mean Väisälä frequency within the survey areas revealed available potential energy in winter above 70 cm2 s-2 and in summer below 20 cm2 s-2 (
The geographical and seasonal distribution of eddy activity has been studied by Väli et al. (2024) using the probability distribution of the Rossby number. Ranges with absolute values in [0, 0.25], [0.25, 0.5], [0.5, 1.0], and [1.0, …] were selected to indicate no eddy activity, weak, moderate, and strong (submesoscale/ageostrophic) eddy activity, respectively. Figure 12 depicts the distribution of weak and moderate eddy activity over the whole study period of 2010–2018. While weak eddy activity represents a linear regime of vortexes with low impact on mixing, moderate eddy activity corresponds to the significant non-linear effects in vortex motions, approaching the ageostrophic submesoscale regime, which may be accompanied by curved threads at Ro = 1. High values of eddy activity are found in the areas of high temporal variability of salinity (Figure 6B). In addition to the frontal regions between the basins and the Western Baltic transition area, high eddy activity is also found in the deep-water path from Stolpe Channel to the Gdansk Basin (Figure 5). Another eddy-active region is along the south-westward brackish water pathways from the Gulf of Finland and the Bothnian Sea along the Swedish coast of the Baltic Proper. The Gulf of Finland is entirely an eddy-active region. Supposedly, high lateral salinity and density gradients favor the generation of eddies due to instabilities; on the other hand, the eddies mix the water masses and reduce the gradients. Although it has been noted that eddies and filaments are often formed during the upwelling processes (Figure 7), the geographical distribution of upwelling occurrence obtained from remote sensing and modelled data (Lehmann et al., 2012) is not directly reflected in the distribution of the Rossby number.
FIGURE 12

Spatial distributions of the occurrence of the weak (Rossby number ranges 0.25 < |Ro| <0.5, (A)) and moderate 0.5 < |Ro| <1, (B) eddy activity in the surface layer for the period 2020–2018. Modified from Väli et al. (2024).
Wind forcing increases the kinetic energy of the surface layer and the whole water column; however, the activated mesoscale and submesoscale motions lag behind the wind speed variations (Väli et al., 2024). In the mechanistic models of coupled basins, wind energy transfer and vertical mixing are governed by turbulence models based on balances of turbulent kinetic energy and its dissipation. The mixed layer dynamics are usually properly simulated in these models. Still, in the deeper layer, penetration of direct wind forcing tends to be too small to explain the observed changes in stratification. A common approach to overcome too low mixing is to include additional mixing terms in the turbulence coefficient, due to the breaking of internal waves. According to Stigebrandt (1987), the extra term is proportional to the inverse of the Brunt-Väisälä frequency. As an example, simulating the end of the enhanced stagnation period 1985–1994, with only a few deep inflows (Figure 13A),
FIGURE 13

Observed salinities from the central Baltic Sea (A) and calculated salinities (B) by adding internal wave energy and Langmuir circulation. Adopted from
There are already many examples of how submesoscale features, generated by a wide range of instability processes at the surface, in the pycnoclines, and in the bottom boundary layer (McWilliams, 2019), participate in the energy cascade from wind and thermohaline forcing to microscale turbulence. However, this research field is far from knowledge “saturation” and further studies are needed.
7 Discussion
The mechanistic models of connected sub-basins give realistic results without resolving the internal sub-basin structures. The mechanistic models resolve the vertical dimension, identify the connection between the sub-basins through the straits, and apply simplified strait flow models. The energy flow from atmospheric forcing ends up in the deeper layers, which are parameterized based on observations during stagnation periods. From model simulations, the Baltic Sea seems to be a strongly forced and damped system. Through winds, heat fluxes, saline water inflow, and river runoff, boundary layer forcing generates currents and eddies and is damped through friction. A well-known observation is, for example, the inertia oscillations that are damped within some oscillation periods (
Turbulent flow involves a multitude of eddies at various scales, some on different scales. This was already illustrated in a drawing by Leonardo da Vinci (1,507–1,509) and described in the book by
The relevant time scale for the mechanistic models is the time it takes to fill a cascade of sub-basins, i.e., the ratio between the volume and the amount of in- or outflow. The propagation time of saline water entering after the Major Baltic Inflow was observed in different deep basins for up to a year (Liblik et al., 2018). The time scale for eddies and submesoscale intrusions along the isopycnals is generally much less, typically 10 days or less.
The mechanistic models assume that the subbasins are horizontally quasi-homogeneous. It means that if the actual contrasts of tracers (e.g., temperature, salinity) within the basin arise (including between the coastal and offshore regions, Figures 7, 11), they are smeared out by meso- and submesoscale processes by a time scale of about 1 month which is a reasonable time frame of the well-working box models. At the same time, the tracer contrasts between the subbasins are governed by meso- and submesoscale frontogenesis, restoring the gradients against mixing. Meandering fronts contribute to the water and tracer transport by shedding the mesoscale eddies and intrusive submesoscale threads. There are indications that meso- and submesoscale mixing contributes more than 90% of the actual lateral mixing. Still, effects of “negative” viscosity and diffusivity also may occur, i.e., during the formation of jet currents and restratification.
Recent years have witnessed a rapid development of ocean descriptions based on machine learning (ML) approaches (
There are examples of combining machine learning with physics-based numerical modeling (Sonnewald et al., 2021;
Future research directions in modelling are expected to enforce true submesoscale-resolving models capable of decadal and centennial model runs, with applied improvements for the non-hydrostatic vertical momentum equation and improved parametrizations from sub-grid scale processes, as well as waves and ice, incorporating the knowledge from observations through machine learning. Longer time scales could be covered by the models of reduced complexity tuned against high-resolution data, either by numerical models of lower resolution or the mechanistic models integrated over the basins.
The large differences in the time scale of basin filling (about 1 year), basin isopycnal through-mixing (>1 month), and eddies (<10 days) illustrate that it is reasonable to assume horizontally homogeneous sub-basins when working on longer time scales with support from the approaches used in the Knudsen theorem. Joint use of the different methods involved in the coupled basin mechanistic models and eddy-resolving models could help understanding the Baltic Sea Earth system, for example, when calculating the ensemble means during climate change studies (Meier et al., 2018).
8 Summary and conclusion
In the present review, we have considered the water exchange in the Baltic Sea based on a historical view of research approaches from basin scales to submesoscale. Mechanistic models of connected sub-basins have been applied in a series of studies starting from the Knudsen theorem. In this class of models, the basin and sub-basin structure was assumed to be horizontally homogeneous, a reasonable assumption on the climate time scale. In parallel, many studies were devoted to mesoscale and submesoscale eddy structures that are highly variable in time and space. These eddy structures are a natural part of the ocean and coastal seas, transforming the large-scale energy through a cascade into smaller scales.
The conclusions could be summarized as follows:
• The mechanistic models and three-dimensional submesoscale approaches complement each other.
• The mechanistic models are useful for interpreting large-scale effects of submesoscale processes; they also allow more numerical experiments and longer modeling periods for climate and long-term environmental studies than three-dimensional eddy-resolving models.
• The submesoscale approaches may guide parametrizations of exchange between the sub-basins and within them.
• Recent submesoscale studies suggest localized eddy-rich regions: Arkona Basin, Gulf of Finland, Irbe Strait, Åland Sea connections, and several coastal areas.
In the coming Baltic Earth phase 2.0, several questions still need new research efforts. A better understanding of the flow of mechanical energy from large-scale forces is needed through basin scale, mesoscale, submesoscale, and microscale. These studies are necessary for the many applied aspects, like offshore building of wind farms and other building activities that may reduce the energy flow, as well as for scenario studies related to other man-made activities under a changing climate.
Statements
Author contributions
JE: Writing – review and editing, Writing – original draft, Investigation, Conceptualization, Methodology, Project administration, Formal Analysis. AO: Writing – review and editing, Methodology, Formal Analysis, Investigation, Writing – original draft, Conceptualization.
Funding
The author(s) declare that financial support was received for the research and/or publication of this article. This work was co-funded by the European Union and the Estonian Research Council via the project TEM-TA38 (Digital Twin of Marine Renewable Energy).
Acknowledgments
This work is part of the Baltic Earth Working Group on Philosophical Views of Baltic Earth and Science. The first version of this paper was partly presented during the 5th Baltic Earth Conference, Jūrmala, Latvia, from 13 May to 17 May 2024, and partly during the Special Baltic Earth Colloquium – achievements, thanks and future challenges, Hamburg, Germany, 4 February 2025. We want to thank the organizer for these two meetings, particularly Marcus Reckermann, for his long service within the Baltex/Baltic Earth program.
Conflict of interest
The authors declare that the research was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest.
Generative AI statement
The author(s) declare that no Generative AI was used in the creation of this manuscript.
Publisher’s note
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Summary
Keywords
Baltic Sea, water budget, water exchange, models, fronts, eddies, submesoscale
Citation
Elken J and Omstedt A (2025) Water exchange in the Baltic Sea: a historical view of research approaches from basin scales to submesoscale. Front. Earth Sci. 13:1598983. doi: 10.3389/feart.2025.1598983
Received
24 March 2025
Accepted
17 June 2025
Published
08 July 2025
Volume
13 - 2025
Edited by
Laura Tuomi, Finnish Meteorological Institute, Finland
Reviewed by
Jaromir Jakacki, Polish Academy of Sciences, Poland
Ulrike Löptien, University of Kiel, Germany
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© 2025 Elken and Omstedt.
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*Correspondence: Jüri Elken, juri.elken@taltech.ee
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