Your new experience awaits. Try the new design now and help us make it even better

ORIGINAL RESEARCH article

Front. Plant Sci., 19 September 2025

Sec. Functional Plant Ecology

Volume 16 - 2025 | https://doi.org/10.3389/fpls.2025.1668310

This article is part of the Research TopicIntegrating Nature-based Solutions for Land Degradation Neutrality and Deriving Co-BenefitsView all 10 articles

The hilly-gully watershed exhibits distinct deep soil moisture characteristics: a comparative study of paired watersheds in the Chinese Loess Plateau

Hongsheng Zhu,Hongsheng Zhu1,2Zihan Wang,Zihan Wang1,2Jiongchang Zhao,Jiongchang Zhao1,2Jiaming LinJiaming Lin1Shuo QianShuo Qian3Liping Wang,Liping Wang1,2Yang Yu,*Yang Yu1,2*Marco CavalliMarco Cavalli4
  • 1School of Soil and Water Conservation, Beijing Forestry University, Beijing, China
  • 2Jixian National Forest Ecosystem Observation and Research Station, Chinese National Ecosystem Research Network (CNERN), School of Soil and Water Conservation, Beijing Forestry University, Beijing, China
  • 3College of Biological Sciences, University of Minnesota, Minneapolis, MN, United States
  • 4National Research Council of Italy, Research Institute for Geo-Hydrological Protection, Padova, Italy

Deep soil moisture constitutes a critical component of hydrological processes in the Loess Plateau, playing an essential role in sustaining vegetation growth, maintaining ecosystem stability, and serving as an important indicator of regional water resource carrying capacity. However, the mechanisms by which long-term vegetation restoration influences deep soil moisture remain insufficiently understood. In this study, we selected two typical paired small watersheds–an Artificial forest watershed and a Farmland watershed-located in the Caijiachuan watershed in the hilly-gully region of the Loess Plateau in western Shanxi Province, China. Based on in situ measurements of soil moisture (0–500 cm) during the 2024 growing season (May-October), the vertical distribution and spatial variability of soil moisture have been systematically analyzed, and the impacts of long-term vegetation restoration on deep soil moisture content have been assessed. The results revealed significant differences between the paired watersheds. The average soil moisture content in the Farmland watershed (0.096 g/g) was significantly higher than in the Artificial forest watershed (0.070 g/g), indicating that artificially introduced vegetation has substantially reduced deep soil moisture reserves. Land use has pronouncedly influenced deep soil moisture, with farmland and native grassland exhibiting the highest moisture retention capacity, while vegetation restoration sites showed the lowest levels. Deep-rooted plantations in the Artificial forest watershed markedly intensified soil moisture deficits in the 200–500 cm layers, whereas the Farmland watershed exhibited comparatively moderate deficits. Moreover, soil moisture spatial heterogeneity was significantly greater in the Farmland watershed, while long-term vegetation restoration promoted a more homogeneous distribution of deep soil moisture. Overall, large-scale restoration dominated by deep-rooted species exerted substantial impacts on deep soil moisture dynamics. These findings provide a scientific basis for vegetation restoration planning and watershed management in the Loess Plateau region.

1 Introduction

Soil moisture constitutes a vital element of the terrestrial system, serving an integrative function in surface processes, particularly within water-limited regions. In these areas, the availability of soil moisture is regarded as the principal determinant of the land productivity and sustainability of watershed management (Gao et al., 2014). Soil moisture demonstrates significant heterogeneity in both spatial and temporal dimensions, even within small watersheds (Dymond et al., 2021; Wang et al., 2019). The characterization of soil moisture variations across diverse spatial and temporal scales is crucial for theoretical understanding and practical applications related to surface runoff and erosion, agriculture management, and ecological restoration (Chatterjee et al., 2022; de Queiroz et al., 2020; Deng et al., 2016).

Soil moisture presents different characteristics in time and space. Numerous studies indicate that soil moisture is affected by a range of environmental factors, including land use, vegetation types, topographical conditions, and soil properties (Fathololoumi et al., 2021; Xu et al., 2021; Yu et al., 2018, 2019). For example, at the hillslope scale, topographic conditions such as slope position and aspect significantly affect soil moisture distribution, with shady slopes and lower positions generally exhibit higher moisture levels due to reduced evaporation and runoff accumulation (Dong et al., 2018); at the watershed scale, land use type, soil properties, and topographical conditions are dominant factors shaping soil moisture variability, where grassland and farmland exhibit differing retention capacities due to root structure and infiltration rates (Huang et al., 2016; Shi et al., 2012); at the regional scale, climatic factors such as precipitation and evapotranspiration predominantly govern soil moisture dynamics, directly influencing moisture content, spatial variability, seasonal fluctuations, and interannual variability (Wang et al., 2012).

Although soil properties and topography tend to remain constant in the short term, land use and climate factors represent the principal dynamic variables (Montenegro and Ragab, 2012; Sharma et al., 2022). On one hand, alterations in land use can significantly disrupt the surface water balance, influencing the allocation of precipitation among evapotranspiration, runoff, and groundwater flow. This, in turn, plays a critical role in determining the distribution of soil moisture within ecosystems. On the other hand, deep soil moisture is less sensitive to short-term climatic fluctuations and land surface changes, but is more closely linked to long-term patterns of infiltration and groundwater dynamics. Studies have shown that deep-layer moisture is predominantly governed by soil texture, soil thickness, topographic wetness index, and antecedent precipitation, with vegetation root depth also contributing significantly to the vertical redistribution of water (Li et al., 2021; Yu et al., 2019). Unlike surface moisture, which responds rapidly to rainfall and evapotranspiration, deep soil moisture content reflects integrated hydrological processes over longer timescales and thus serves as a more stable indicator of ecohydrological conditions (Wang et al., 2024a, 2024).

The Loess Plateau is an environmentally sensitive area, soil loss and water erosion are the main environmental problems in this area. Since 1999, the “Grain to Green Program” has promoted introduced vegetation to combat soil erosion, vegetation restoration proving to be the most effective measure in ecological management (Bai et al., 2024; Wang et al., 2021; Yu et al., 2020). Despite significant progress in ecological engineering, large-scale revegetation has led to excessive water consumption, resulting in dried soil layers and stunted old trees, thus weakening the expected effect of afforestation in reducing water loss and improving the local environment (Mei and Ma, 2022; Yao et al., 2016; Yu et al., 2024). Deep soil moisture is difficult to replenish. A decline in soil moisture has many other consequences for local water resources. For instance, Wang et al. (2024) found that in China’s monsoon loess critical zone, intensive land-use conversions since 1999 have been the predominant catalyst of deep soil moisture decline, exceeding the effects of climate change and threatening soil moisture security across the Loess Plateau. Cheng et al. (2020) reported that in the semi-arid regions of northern China, vegetation restoration has significantly improved precipitation utilization. However, it has inhibited the recharge of deep soil moisture. Jia and Shao, 2014 emphasized that vegetation type significantly affects the dynamics of deep soil moisture. In addition to the increasing potential evapotranspiration, plant water consumption has substantially degraded soil moisture conditions in the experimental plots.

Soil moisture monitoring at the watershed scale is a crucial component in understanding the interactions between water and vegetation. Recent research on small watersheds has predominantly concentrated on slopes or individual land-use types. However, there is a paucity of studies examining the impact of vegetation restoration on deep soil moisture at the watershed scale. The implementation of paired watershed monitoring is a primary method for elucidating hydrological processes; nevertheless, few studies have systematically examined deep water dynamics paired with watersheds. In this study, a paired-site methodology has been employed to assess the variation and magnitude of soil moisture resulting from long-term vegetation restoration in the hilly-gully region of the Loess Plateau, China. The main research purposes of this study were (i) to compare the differences of soil moisture variation across layers between paired watersheds; (ii) to explore the relationship between land uses and soil moisture variability, and (iii) to examine the effects of long-term vegetation restoration on deep soil moisture distribution.

2 Materials and methods

2.1 Study area

The study area is located in the Caijiachuan watershed in the Loess Plateau (Figure 1) in the western part of Shanxi Province, China, with geographical coordinates ranging from 110°39′45″ to 110°47′45″E and 36°14′27″ to 36°18′23″N. Since afforestation began in 1990, the forest coverage rate in the watershed has exceeded 80%, resulting in the establishment of approximately 1,000 hectares of soil and water conservation forest. The tree layer is dominated by Robinia pseudoacacia (RP) and Pinus tabuliformis (PT), the shrub layer by Rosa xanthina (RX), and the herbaceous layer by Festuca elata (FE). The Artificial forest watershed and the Farmland watershed are two paired watersheds affiliated with the Caijiachuan watershed. Prior to vegetation restoration, the Artificial forest watershed was mainly farmland and shared similar land-use characteristics with the current Farmland watershed, which is geographically adjacent and thus selected as a reference (paired) watershed. Both watersheds are located in a typical loess residual plateau gully region, characterized by brown soils derived from loess parent material. The area experiences a warm-temperate continental climate, with rainfall primarily concentrated between June and September. The annual sunshine duration is approximately 2,563 hours, the average annual precipitation is 579 mm, and the average potential evapotranspiration is 1,723.9 mm. Rainfall patterns exhibit a uniform spatial distribution across the two watersheds. The Artificial forest watershed is situated in the central part of the Caijiachuan watershed, with an elevation ranging from 969 to 1,232 m and a total area of 1.52 km². It features high terrain in the northwest and low terrain in the southeast, with a general north-south orientation. Land use is dominated by vegetation restoration species (RP, PT, RX). In contrast, the Farmland watershed lies in the eastern part of the Caijiachuan watershed, with an elevation ranging from 915 to 1,110 m and an area of 0.76 km². Its terrain is high in the southeast and low in the northwest, also aligned in a north-south direction. The dominant land-use types are rain-fed farmland planted with Zea mays (ZM) and economic forest with Malus domestica (MD). The introduced vegetation types in the Caijiachuan watershed include RP, PT, and RX.

Figure 1
Elevation and land use maps depict the Loess Plateau and Caijiachuan watershed, showcasing varied elevations in meters. The top image shows the broader Loess Plateau, while the middle highlights the watershed. The bottom section details land use types like vegetated restoration land, economic forestland, native grassland, and farmland in two watersheds, labeled with elevation ranges.

Figure 1. Location of the study area and experimental sites in two watersheds.

2.2 Field monitoring and experimental design

In the Artificial forest watershed and the Farmland watershed, there are different land use types. This study focused on 4 different land use types with different dominant plant species on them, which are native grassland (Festuca elata, FE), vegetated restoration land (Robinia pseudoacacia, RP; Pinus tabuliformis, PT; Rosa xanthina, RX), economic forestland (Malus domestica, MD), and farmland (Zea mays, ZM). In the Artificial Forest watershed, there were three land use types: native grassland, vegetated restoration land, and farmland, and in the Farmland watershed, there are also three land use types, which are vegetated restoration land, farmland, and economic forestland. Considering that the soil properties of the Artificial forest watershed and the Farmland watershed are generally consistent, this study, based on topographic characteristics and vegetation distribution, has established 34 experimental sites in the Artificial forest watershed: 30 of them were located within vegetated restoration land (including 19 in RP, 3 in PT, and 8 in RX), 3 of them were in native grassland (FE), and 1 of them were in farmland (ZM) (Table 1), and 22 experimental sites in the Farmland watershed: 8 of them were in vegetated restoration land (including 3 in RP and 5 in RX), 6 of them were in economic forestland (MD), and another 8 of them were in farmland (ZM) (Table 2), in August 2024 using a grid-based sampling design. At each experimental site, soil samples were collected with a soil auger to measure the gravimetric soil moisture (g/g) within the 0–500 cm soil profile. Given the pronounced topographic variability and the complex distribution of vegetation types across the Loess Plateau, the grid-based layout was adopted to enhance the spatial representativeness of the experimental sites, facilitate comparisons of soil moisture status among different land use types, and provide a robust foundation for subsequent modeling of spatial variability in deep soil moisture. During the field surveys, slope gradient, slope aspect, vegetation type, and vegetation cover were specifically documented at each experimental site. In addition, a handheld GPS device was used to measure and record the latitude, longitude, and elevation of each site.

Table 1
www.frontiersin.org

Table 1. Topographic and vegetation characteristics of monitoring plots in the Artificial forest watershed.

Table 2
www.frontiersin.org

Table 2. Topographic and vegetation characteristics of monitoring plots in the Farmland watershed.

2.3 Data collection

In the growing season of 2024 (May to October), soil moisture content in the 0–500 cm layer was measured at each experimental site. Three sampling profiles were randomly selected per site to obtain the average soil moisture content. Soil samples were collected with a 5 cm diameter auger at 10 cm intervals for the 0–100 cm layer and 20 cm intervals for the 100–500 cm layer, each profile yielded 30 samples, generating a total collection of 5040 samples across all study sites. According to previous studies conducted in the Loess Plateau (Yu et al., 2019), the annual precipitation infiltration depth generally does not exceed 200 cm, and soil moisture below this depth exhibits relatively small fluctuations and remains comparatively stable. Therefore, in this study, the 0–200 cm layer was defined as shallow soil, while the 200–500 cm layer was defined as deep soil. Samples were immediately sealed in aluminum cylinders after collection and weighed. They were then dried in an oven at 105°C until a constant weight was achieved. Soil moisture content (g/g) was calculated as the ratio of the mass lost during oven drying to the constant dry weight.

To analyze the spatial distribution of soil moisture, the Inverse Distance Weighting (IDW) method was used to interpolate soil moisture content and generate spatial distribution maps at different depths for both watersheds. ArcGIS software was employed for IDW interpolation and mapping, and its spatial analysis tools were used to perform basic statistical analyses based on the interpolated data.

2.4 Statistical analysis

The soil moisture content and profile distribution in each watershed was calculated by taking the average of all experimental sites in each soil layer. The depth-averaged gravimetric soil moisture content θij (g/g) at each experimental site was calculated by Equation 1.

θij=1ii=1iθi(1)

where I is the number of measured strata at site j and θij is the soil moisture content of stratum i calculated from three randomly sampled profiles. In the following analysis, gravimetric soil moisture content is calculated on a per-100-centimeter basis, with I = 5.

The depth-averaged gravimetric soil moisture content θm (g/g) for each land use type was calculated by Equation 2.

θm=1kk=1kθij(2)

where k is the number of experimental sites for each land use type.

The depth-averaged volumetric soil moisture content θv (%) for each land use type was calculated by Equation 3.

θv=θmi×ρbi(3)

where ρbi is the average soil bulk density of layer i for each land use type.

The soil moisture content depletion degree (SMCD) (%) was calculated by Equation 4.

SMCD=1kk=1kθvlθvcθvl(4)

where θvl represents the volumetric soil moisture content of the layer i of farmland (Zea mays, ZM), and θvc represents the volumetric soil moisture content of the layer i under different vegetation types.

This study utilized the Inverse Distance Weighting (IDW) method within ArcGIS 10.3 to interpolate and analyze the spatial distribution of soil moisture in the Artificial Forest Watershed and the Farmland Watershed (Gao and Yang, 2023). Spatial distribution maps of soil moisture were generated using this method. IDW estimates the values of unsampled locations through a linear weighting of known sample sites, where the weights are inversely proportional to the distance between the sample and the target sites. The IDW method was calculated by Equation 5.

Z^(u)=i=1nZid(u,ui)Pi=1n1d(u,ui)P(5)

where Z^(u) is the estimated value at the unknown site u; Zi is the observed value at the known site ui; d(u,ui) represents the distance between the unknown site u and the known site ui; and P is the power parameter that controls the influence of distance on the weighting (Pal and Maity, 2021).

Basic descriptive statistics, including the mean, standard deviation (SD), and coefficient of variation (CV), were calculated for each measurement. Prior to conducting the paired t-test and one-way analysis of variance (ANOVA), the Shapiro–Wilk test was applied to assess the normality of the data. Variables that did not meet the normality assumption were log-transformed before statistical analysis. A paired t-test was utilized to examine the differences in soil moisture across soil layers between the two watersheds. To assess the impact of various land use types on the overall variability of soil moisture, a one-way analysis of variance (ANOVA) was conducted. Post hoc multiple comparisons were executed using the Least Significant Difference (LSD) method. Microsoft Excel was employed for preliminary data organization, whereas SPSS and Origin were used for advanced statistical analysis and data visualization.

3 Results

3.1 Vertical soil moisture variation

As shown in Figure 2, a significant difference has occurred in soil moisture content between the two watersheds (p < 0.05). Soil moisture in the Artificial forest watershed ranged from 0.038 to 0.082 g/g, whereas in the Farmland watershed, it ranged from 0.082 to 0.130 g/g, the overall soil moisture content in the Farmland watershed is higher than that in the Artificial forest watershed. In terms of vertical distribution, the average soil moisture content in the Artificial forest watershed gradually increased with soil depth before stabilizing, whereas in the Farmland watershed, it originally decreased, then increased, and eventually stabilized. The standard deviation of soil moisture in the Farmland watershed was greater than in the Artificial forest watershed.

Figure 2
Graph depicting soil moisture content (g/g) versus soil depth (cm) with two lines: red squares for artificial forest watershed and blue circles for farmland watershed. Error bars are included.

Figure 2. Profile distribution of the mean soil moisture content in two watersheds, the error bars indicate standard deviation.

As shown in Figure 3, there was a significant difference (p < 0.05) in the average soil moisture content across different soil layers between the Artificial forest watershed and the Farmland watershed. Specifically, in the 0–100 cm soil layer, the average soil moisture content in the Farmland watershed was 0.105 g/g, which was 72.13% higher than in the Artificial forest watershed (0.061 g/g). In the 100–200 cm soil layer, soil moisture in the Farmland watershed was 0.085 g/g, 16.44% higher than in the Artificial forest watershed (0.073 g/g). In the 200–300 cm soil layer, the Farmland watershed recorded 0.092 g/g, 24.32% higher than the Artificial forest watershed (0.074 g/g). In the 300–400 cm soil layer, the moisture content in the Farmland watershed was 0.096 g/g, 29.73% higher than in the Artificial forest watershed (0.074 g/g). In the 400–500 cm soil layer, it was 0.094 g/g in the Farmland watershed, 20.51% higher than in the Artificial forest watershed (0.078 g/g). Overall, the average soil moisture content in the 0–500 cm layer in the Farmland watershed was 0.096 g/g, representing a 37.14% increase compared to the Artificial forest watershed (0.070 g/g).

Figure 3
Bar graph showing soil moisture content across different depths (0-100 cm, 100-200 cm, 200-300 cm, 300-400 cm, 400-500 cm, 0-500 cm) for artificial forest and farmland watersheds. Farmland consistently has higher moisture content than the artificial forest.

Figure 3. Comparison of soil moisture content of different soil layers in two watersheds, error bars indicate standard deviation, different uppercase letters above the bars indicate significant differences in soil moisture content between the two watersheds.

3.2 Soil moisture of different land use types in paired watersheds

In the two watersheds, soil moisture content was highest in farmland (ZM), followed by native grassland (FE), and lower in the vegetated restoration land (RP, PT, RX), and economic forestland (MD) (Figure 4; Table 3). After the LSD test (Table 3), the differences in soil moisture between different vegetation types in the same watershed were statistically significant (p < 0.05).

Figure 4
Two line graphs depict soil moisture content (g/g) versus soil depth (cm). Graph A shows five lines labeled FE, RP, PT, RX, and ZM. Graph B shows lines RX, RP, MD, and ZM. Each line represents different data series, with varying trends in moisture content up to 500 cm depth. The legend indicates different colored and shaped markers for each series.

Figure 4. Soil moisture in different land use types in two watersheds (A) Artificial forest watershed and (B) Farmland watershed. FE, native grassland (Festuca elata); RP, PT, and RX, vegetated restoration land (Robinia pseudoacacia, Pinus tabuliformis, and Rosa xanthina, respectively); ZM, farmland (Zea mays); MD, economic forestland (Malus domestica).

Table 3
www.frontiersin.org

Table 3. Soil moisture (g/g) of 0–500 cm soil layers in different land use types.

In the Artificial forest watershed, there was no significant difference in mean soil moisture content among RP, PT, and RX in the vegetated restoration land (p > 0.05), in the 0–200 cm soil layer, the mean soil moisture content was significantly higher in native grassland (FE) than in the vegetated restoration land (RP, PT, RX) and farmland (ZM) (p < 0.05), and in the 200–500 cm soil layer, the mean soil moisture content was significantly higher in both native grassland (FE) and farmland (ZM) than in the vegetated restoration land (RP, PT, RX) (p < 0.05). In the Artificial forest watershed, the average soil moisture content in the 0–500 cm soil layer of different vegetation types, in descending order, was farmland (ZM) (0.099 g/g), native grassland (FE) (0.091 g/g), vegetated restoration land (RX) (0.071 g/g), vegetated restoration land (PT) (0.070 g/g), and vegetated restoration land (RP) (0.068 g/g), with farmland (ZM) and native grassland (FE) all having significantly higher soil moisture content than vegetated restoration land (RP, PT, RX).

In contrast, in the 0–100 cm soil layer of Farmland watershed, the mean soil moisture content was significantly higher (p < 0.05) in farmland (ZM) and vegetated restoration land (RP) than in vegetated restoration land (RX) and economic forestland (MD), with no significant differences detected between economic forestland (MD)-vegetated restoration land (RP) pair and vegetated restoration land (RX)-economic forestland (MD) pair (p > 0.05), and in the 100–500 cm soil layer, the mean soil moisture content was significantly higher (p < 0.05) in farmland (ZM) than in vegetated restoration land (RP, RX), and economic forestland (MD). No significant differences have occurred in vegetated restoration land (RP), and statistical analysis revealed no detectable differences (p > 0.05) between the soil moisture of vegetated restoration land (RX) and economic forestland (MD). In the Farmland watershed, across the 0–500 cm profile, farmland (ZM) (0.120 g/g) maintained the highest average soil moisture content, followed in decreasing order by vegetated restoration land (RP) (0.094 g/g), economic forestland (MD) (0.084 g/g), and vegetated restoration land (RX) (0.080 g/g), in which the average soil moisture content of farmland (ZM) was significantly higher than vegetated restoration land (RP) (p < 0.05), the average soil moisture content of vegetated restoration land (RP) were significantly higher than vegetated restoration land (RX) and economic forestland (MD) (p < 0.05), and there was no significant difference between vegetated restoration land (RX) and economic forestland (MD) (p > 0.05).

3.3 Soil moisture depletion of different land use types in paired watersheds

Compared to farmland (ZM), both Artificial forest watershed and Farmland watershed exhibited varying degrees of soil moisture deficit (SMCD), with the deficit being more severe in Artificial forest watershed (Figure 5, Table 4).

Figure 5
Bar graphs A and B depict SMCD percentages across soil layers from zero to five hundred centimeters. Graph A includes four color-coded bars: pink (FE), blue (RP), red (PT), teal (RX). Graph B has three bars: teal (RX), blue (RP), brown (MD). Both graphs show varied SMCD levels across soil layers with labeled error bars.

Figure 5. Variations in soil moisture depletion degree (SMCD) across soil layers under different land use types in two watersheds: (A) Artificial forest watershed and (B) Farmland watershed. Different lowercase and uppercase letters above the bars indicate significant differences between different land uses at the same soil depth and between the same land use at different soil depths (p < 0.05). FE, native grassland (Festuca elata); RP, PT, and RX, vegetated restoration land (Robinia pseudoacacia, Pinus tabuliformis, and Rosa xanthina, respectively); MD, economic forestland (Malus domestica).

Table 4
www.frontiersin.org

Table 4. Soil moisture depletion degree (SMCD) of the 0–500 cm soil layers.

Within the Artificial forest watershed, significant differences in SMCD were observed across soil layers for the same vegetation type. In native grassland (FE), the 0–100 cm and 100–200 cm layers showed no moisture deficit, while the 200–300 cm, 300–400 cm, and 400–500 cm layers exhibited deficits of 26.68%, 36.23%, and 25.10%, respectively. Similarly, no deficit was found in the 0–100 cm layer of vegetated restoration land (RP), but the 100–200 cm, 200–300 cm, 300–400 cm, and 400–500 cm layers showed SMCDs of 17.74%, 45.16%, 47.75%, and 42.03%, respectively. SMCD values were significantly higher in the 200–500 cm layers than in the 100–200 cm layer. For vegetated restoration land (PT and RX), all soil layers exhibited moisture deficits. In vegetated restoration land (PT), SMCD values reached 42.33% in the 200–300 cm, 300–400 cm, and 400–500 cm layers, which were significantly higher than the values in the 0–100 cm (6.60%) and 100–200 cm (17.79%) layers. Similarly, vegetated restoration land (RX) showed SMCDs of 43.48%, 44.92%, and 37.89% in the 200–300 cm, 300–400 cm, and 400–500 cm layers, all significantly higher than those in the 0–100 cm (9.95%) and 100–200 cm (13.77%) layers. Additionally, SMCD varied significantly among different vegetation types within the same soil layer. The vegetated restoration land (RP, PT, RX) exhibited significantly higher SMCD values than native grassland (FE) across all layers from 0–500 cm.

Within the Farmland watershed, the soil moisture content difference (SMCD) of vegetated restoration land (RX) and economic forestland (MD) did not differ significantly among soil layers. Vegetated restoration land (RP) did not experience water deficit in the 0–100 cm soil layer; however, soil moisture depletion was observed in the 100–200 cm, 200–300 cm, 300–400 cm, and 400–500 cm layers, with SMCDs of 11.35%, 14.11%, 12.48%, and 17.39%, respectively. In addition, significant differences in SMCD were observed among different vegetation types within the same soil layer, with vegetated restoration land (RX) and economic forestland (MD) exhibiting significantly higher SMCD values than vegetated restoration land (RP) across all soil layers from 0–500 cm.

3.4 Spatial distribution of soil moisture in different depths

Figure 6 was generated with the Inverse Distance Weighting (IDW) method to characterize the spatial distribution of soil moisture in the Artificial forest watershed and the Farmland watershed. The results indicate that soil moisture distribution was closely associated with land use type, with relatively higher soil moisture in farmland (ZM) and native grassland (FE), and lower moisture levels at vegetated restoration land (RP, PT, RX). In the Farmland watershed, soil moisture showed a spatial pattern of high values in the northwest and low values in the southeast, while the Artificial forest watershed exhibited the opposite trend-lower values in the northwest and higher values in the southeast. The maximum mean soil moisture content in the 0–500 cm layer of the Artificial forest watershed occurred in native grassland (FE) (0.101 g/g at site 24), while the minimum occurred in vegetated restoration land (RP) (0.0526 g/g at site 6). In the Farmland watershed, the highest mean soil moisture was observed in farmland (ZM) (0.170 g/g at site 5), and the lowest in vegetated restoration land (RX) (0.067 g/g at site 14).

Figure 6
Maps labeled A to E showing soil moisture content within watershed boundaries, using grayscale to depict varying moisture levels from 0.055 to 0.195 grams per gram. Each map is divided with darker areas indicating higher moisture content. A legend explains the grayscale values and a scale bar denotes distance in kilometers.

Figure 6. Soil moisture content interpolated by the IDW method in different soil depths. (A) 0–100 cm, (B) 100–200 cm, (C) 200–300 cm, (D) 300–400 cm, and (E) 400–500 cm.

Table 5 presents the statistics of soil moisture content across different soil layers in the two watersheds. In the 0–500 cm depth range, the coefficient of variation in the Artificial forest watershed ranges from 19.98% to 25.72%, with a standard deviation of 0.013–0.017. In the Farmland watershed, the coefficient of variation ranges from 29.95% to 31.29%, and the standard deviation ranges from 0.026 to 0.031. Both the coefficient of variation and standard deviation in the Farmland watershed are higher than those in the Artificial forest watershed.

Table 5
www.frontiersin.org

Table 5. Basic statistics on soil moisture content (g/g) in two watersheds based on IDW.

4 Discussion

4.1 Vertical distribution characteristics of soil moisture

The monitoring of deep soil moisture revealed that there were significant differences in soil moisture content between the paired watersheds (Figure 2). The Farmland watershed exhibited a markedly higher average soil moisture compared to the Artificial forest watershed. This phenomenon is consistent with the findings of Jia et al. (2017) and Liu et al. (2018), both of which indicated that, in typical vegetated restoration areas of the Loess Plateau, prolonged high-density afforestation has resulted in sustained depletion of deep soil moisture. The dynamics of soil moisture are directly influenced by precipitation, topography, and vegetation. Among these factors, precipitation constitutes a critical constraint on vegetated restoration and reconstruction in the Loess Plateau and represents one of the principal sources of soil moisture (Hou et al., 2018; Jin et al., 2018). In this study, the selected paired watersheds are adjacent to each other, with similar elevation and spatial precipitation distribution. Additionally, the Loess Plateau is characterized by deep soil layers, and soil moisture below the infiltration depth in restored vegetation areas remains relatively stable with minimal interannual fluctuations (Yang et al., 2012). Although the analysis in this study is based on monitoring data from a single growing season, short-term climatic variations have limited effects on deep soil moisture. Therefore, the primary factor responsible for the observed differences in deep soil moisture between the paired watersheds is likely the influence of vegetation roots under long-term restoration. The introduced vegetation in the Artificial forest watershed has developed extensive root systems, which, under identical rainfall conditions, exert a greater influence on soil moisture content compared to farmland and grassland (Wyatt et al., 2021). Moreover, the vegetation density in the Artificial forest watershed considerably exceeds the carrying capacity of natural precipitation, and rainfall is insufficient to meet the growth and development demands of such biomass-rich artificial plantations, thereby leading to excessive depletion of soil moisture (Jin et al., 2018). In contrast, the Farmland watershed is mainly cultivated with shallow-rooted annual crops such as ZM, which primarily consume surface soil moisture, allowing deeper moisture to be conserved.

By comparing soil moisture content across different soil depths in the two watersheds (Figure 3), it was found that the soil moisture content of the 0–100 cm layer in the Farmland watershed was 0.105 g/g, which was significantly higher than that in the Artificial forest watershed (0.061 g/g), representing a difference of 72.13%. In the 100–500 cm soil layer, however, the difference in soil moisture content narrowed to 16.44%, 24.32%, 29.73%, and 20.51%. This could be attributed to the fact that the lateral roots of the introduced vegetation species (RP, PT, RX) in the Artificial forest watershed are mainly distributed within the 0–100 cm layer, where considerable amounts of soil moisture are consumed during the growing season to sustain plant development (Cui et al., 2023; Zhou et al., 2022). Additionally, precipitation predominantly affects shallow soil moisture. Due to extensive vegetation cover in the Artificial forest watershed, canopy interception reduces surface infiltration, resulting in lower soil moisture content (O’Donnell et al., 2021; Wu et al., 2021). In the Farmland watershed, long-term cultivation practices have left crop residues and surface organic matter that help reduce bare soil exposure and minimize evaporation losses of surface soil moisture (Tuure et al., 2021). Furthermore, continuous tillage has altered the soil structure, promoted infiltration, and consequently increased soil moisture content in the 0–100 cm layer (Wang et al., 2017). The deep soil moisture content in the Artificial forest watershed remained lower than in the Farmland watershed. This may be because, under long-term vegetation restoration, plants tend to develop deeper roots to access moisture when the upper soil layers experience water stress, thereby depleting deep soil moisture and forming a dried soil layer (Tijdeman and Menzel, 2021; Wang et al., 2024).

In addition to root distribution and vegetation density, species-specific water use characteristics further elucidate the observed contrasts in deep soil moisture. Sap flow and eddy covariance studies in the Loess Plateau have shown that Robinia pseudoacacia (RP) exhibits annual evapotranspiration levels of approximately 480–550 mm·yr−1. Due to its deep root system and high canopy transpiration demand, the overall water consumption of RP often exceeds the mean annual precipitation (Wu et al., 2021). By contrast, Zea mays (ZM) generally has an annual evapotranspiration of 350–450 mm·yr−1, which is more closely aligned with regional precipitation inputs (Liu et al., 2010). This indicates that ZM primarily relies on seasonal rainfall, whereas RP continuously depletes deep soil moisture. These interspecific differences in water use efficiency further demonstrate the importance of considering evapotranspiration demand when selecting restoration species in water-limited regions such as the Loess Plateau.

4.2 Mechanisms driving the spatial variability of deep soil moisture

This study found that the spatial heterogeneity of soil moisture in the Farmland watershed was considerably higher than that in the Artificial forest watershed (Figure 6). Across the 0–500 cm profile, the coefficient of variation in the Farmland watershed reached 31.25%, exceeding the 24.29% observed in the Artificial forest watershed. In all soil layers, the coefficient of variation in the Farmland watershed was consistently higher, which aligns with the findings of Yang et al. (2014), who reported that the distribution of soil moisture in the Farmland watersheds is more complex compared to the Artificial forest watersheds. The primary reason for this difference lies in the contrasting land use structures between the two watersheds. The Artificial forest watershed has undergone long-term vegetation restoration, resulting in a predominantly monocultural vegetation pattern dominated by artificial forest, which leads to relatively uniform soil moisture distribution. By contrast, the Farmland watershed comprises a mosaic of farmland, economic forest, and shrubland, thereby increasing the spatial variability of soil moisture distribution. The significant differences in deep soil moisture between the two watersheds indicate that complex land use structures are associated with higher spatial heterogeneity of deep soil moisture, whereas large-scale vegetation restoration tends to enhance the spatial homogeneity of deep soil moisture.

In addition, soil and water conservation projects also play a crucial role in shaping the spatial pattern of deep soil moisture. Among the 22 experimental sites in the Farmland watershed, sites 1, 5, 8, and 13 exhibited substantially higher soil moisture than other sites. This is likely because these sites were located within check dams, which are typical soil and water conservation measures in the Farmland watersheds. Check dams promote water infiltration, reduce surface runoff, and control soil erosion, thereby contributing to increased soil moisture (Helman and Mussery, 2020; Luan et al., 2022).In the Artificial forest watershed, the highest soil moisture in the 0–200 cm layer was observed at site 24, possibly because this site was situated in native grassland (FE). FE has shallow root systems, resulting in less uptake of deep soil moisture (Ye et al., 2019). Additionally, the canopy height and biomass of FE are relatively low, leading to lower canopy interception and evaporation compared to other vegetation types. As a result, a greater proportion of precipitation infiltrates into the soil, maintaining higher soil moisture content (Jian et al., 2015). In the 200–500 cm layer, the highest soil moisture was recorded at sites 19 and 34. This may be attributed to site 19 being located in farmland (ZM), where tillage practices modify the structure of surface soil aggregates, increase soil porosity, and facilitate infiltration of rainfall, thereby accelerating the percolation of moisture into deeper soil layers. Site 34 was located at the outlet of the watershed, in a relatively low-lying terrain position. Under identical precipitation or snowmelt input conditions, water tends to accumulate along the slope and infiltrate into deep soil layers, forming a moisture convergence zone and resulting in higher deep soil moisture content (Chang et al., 2024).

4.3 Implications for vegetation restoration

In the Loess Plateau region, deep soil moisture plays a critical role in sustaining the growth and ecological functions of artificial vegetation (Figure 4). This study demonstrated that, in the Artificial forest watershed, soil moisture content across all soil layers under vegetated restoration land (RP, PT, RX) was significantly lower compared to farmland (ZM) and native grassland (FE). Specifically, the average soil moisture content (0–500 cm) in the Artificial forest watershed was significantly higher in ZM (0.099 g/g) and FE (0.091 g/g) than in RP (0.068 g/g), PT (0.070 g/g), and RX (0.071 g/g). A similar pattern was observed in the Farmland watershed, where the difference in moisture content between vegetated restoration land and farmland was also significant. In the Farmland watershed, farmland (ZM) showed the highest average moisture content in the 0–500 cm soil layer (0.120 g/g), significantly higher than RP (0.094 g/g), MD (0.080 g/g), and RX (0.080 g/g). This difference may be attributed to the annual crop (ZM), which has a shallow root system and short life cycle, consuming mainly surface soil moisture and thus conserving deeper soil moisture (Shen et al., 2014; Xiao et al., 2014). Native grassland (FE), despite having year-round coverage, also possesses relatively shallow roots that consume less deep-layer moisture, thus maintaining higher deep soil moisture levels (Zhang et al., 2024). In contrast, vegetated restoration sites (RP, PT, RX) have deeper root systems and, over prolonged restoration periods, continuously consume deep soil moisture through transpiration, resulting in a lower soil moisture content (Liu et al., 2024a).

The difference in soil moisture content between vegetated restoration sites and farmland can represent the extent of soil moisture depletion caused by vegetation restoration. Results from this study (Figure 5) indicate that the soil moisture deficit at vegetated restoration sites (RP, PT, and RX) was significantly greater than that of native grassland (FE) within the 0–500 cm soil layer in the Artificial forest watershed. Specifically, in terms of soil depth, the soil moisture content deficit (SMCD) for RP, PT, and RX in the 0–100 cm soil layer was −7.52%, 6.60%, and 9.90%, respectively, while the most severe water depletion occurred in the 300–400 cm soil layer, with deficits of 47.75%, 46.94%, and 44.90%, respectively. This pattern demonstrates that artificial vegetation significantly depleted deep soil moisture reserves, with a more pronounced depletion effect in deeper soil layers, leading to the formation of a deep, dry soil layer that gradually restricts forest growth (Zhang et al., 2018). In contrast, soil moisture depletion by artificial vegetation in the Farmland watershed did not reach such extreme levels. In the Farmland watershed, RP did not exhibit a soil moisture deficit in the 0–100 cm layer, and deeper layers showed only moderate deficits ranging from 11.35% to 17.39%. This moderate deficit may result from the smaller scale and lower density of the RP in the Farmland watershed compared to the Artificial forest watershed, resulting in relatively lower deep soil moisture consumption (Liu et al., 2024b; Zhang et al., 2018). These findings suggest that large-scale vegetation restoration, particularly with deep-rooted species, significantly reduces deep soil moisture content, and that the depletion mechanism is closely related to the water-use characteristics of vegetation. This interpretation is consistent with Zhou et al. (2024), who emphasized that vegetation type exerts a strong influence on deep soil moisture dynamics, with deep-rooted broadleaf species contributing more to deep soil drying, whereas shallow-rooted vegetation maintains a more balanced soil moisture regime. The scarcity of deep soil moisture therefore poses a critical challenge to the sustainability of afforestation and vegetation restoration efforts in water-limited regions. Consequently, it is advisable to avoid large-scale afforestation with deep-rooted monocultures, as this practice may expedite the development of deep dry soil layers. The density of afforestation efforts should be optimized based on rigorous scientific analysis of local hydrological conditions. For existing plantations, adaptive management strategies—such as thinning to decrease stand density or replacing species—should be implemented to mitigate deep soil moisture deficits and promote the long-term sustainability of vegetation restoration efforts.

5 Conclusions

This study has systematically analyzed the vertical distribution characteristics and spatial variability of soil moisture (0–500 cm) in typical paired watersheds (Farmland watershed and Artificial forest watershed) within the Caijiachuan watershed in the hilly-gully region of the Loess Plateau in western Shanxi Province. The results demonstrated that long-term vegetation restoration substantially reduced deep soil moisture reserves. The mean soil moisture content in the Artificial forest watershed (0.070 g/g) was significantly lower than that in the Farmland watershed (0.096 g/g), indicating that artificially introduced deep-rooted vegetation markedly intensified soil moisture deficits in the 200–500 cm layers. Farmland and native grassland exhibited stronger capacities for maintaining deep soil moisture, whereas vegetation restoration sites consistently showed pronounced deep moisture depletion. In addition, the spatial heterogeneity of deep soil moisture was significantly higher in the Farmland watershed, while long-term afforestation promoted a more homogeneous spatial distribution of soil moisture content. Overall, the findings indicate that large-scale vegetation restoration dominated by deep-rooted species can lead to substantial deep soil desiccation, posing potential risks to vegetation sustainability and regional water balance. Therefore, future land-use planning and vegetation restoration efforts in the Loess hilly-gully watershed should carefully balance the trade-offs between ecological restoration and soil moisture availability, scientifically select and arrange appropriate vegetation types, and implement adaptive management strategies to ensure the long-term sustainability of ecosystem functions.

Data availability statement

The raw data supporting the conclusions of this article will be made available by the authors, without undue reservation.

Author contributions

HZ: Data curation, Validation, Visualization, Writing – original draft, Writing – review & editing. YY: Conceptualization, Investigation, Methodology, Supervision, Writing – original draft, Writing – review & editing. ZW: Resources, Validation, Visualization, Investigation, Software, Writing – review & editing. JZ: Investigation, Resources, Visualization, Writing – review & editing. JL: Investigation, Methodology, Writing – review & editing. SQ: Data curation, Investigation, Writing – review & editing. LW: Investigation, Resources, Writing – review & editing. MC: Supervision, Writing – review & editing.

Funding

The author(s) declare financial support was received for the research and/or publication of this article. This study was supported by the National Key Research and Development Program of China (No. 2023YFF1305101), the National Natural Science Foundation of China (42177310), and the Fundamental Research Funds for the Central Universities (QNTD202508).

Conflict of interest

The authors declare that the research was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest.

Generative AI statement

The author(s) declare that no Generative AI was used in the creation of this manuscript.

Any alternative text (alt text) provided alongside figures in this article has been generated by Frontiers with the support of artificial intelligence and reasonable efforts have been made to ensure accuracy, including review by the authors wherever possible. If you identify any issues, please contact us.

Publisher’s note

All claims expressed in this article are solely those of the authors and do not necessarily represent those of their affiliated organizations, or those of the publisher, the editors and the reviewers. Any product that may be evaluated in this article, or claim that may be made by its manufacturer, is not guaranteed or endorsed by the publisher.

References

Bai, R., Wang, X., Li, J., Yang, F., Shangguan, Z., and Deng, L. (2024). The impact of vegetation reconstruction on soil erosion in the loess plateau. J. Environ. Manage. 363, 121382. doi: 10.1016/j.jenvman.2024.121382

PubMed Abstract | Crossref Full Text | Google Scholar

Chang, Y., Han, L., Chen, R., Liu, Z., Li, Y., Zhao, Z., et al. (2024). Modelling of the trade-off between the deep soil moisture and vegetation restoration in the hilly area of the loess plateau, China. J. Hydrology 645, 132274. doi: 10.1016/j.jhydrol.2024.132274

Crossref Full Text | Google Scholar

Chatterjee, S., Desai, A. R., Zhu, J., Townsend, P. A., and Huang, J. (2022). Soil moisture as an essential component for delineating and forecasting agricultural rather than meteorological drought. Remote Sens. Environ. 269, 112833. doi: 10.1016/j.rse.2021.112833

Crossref Full Text | Google Scholar

Cheng, Y., Li, X., Wang, Y., Zhan, H., Yang, W., and Jiang, Q. (2020). New measures of deep soil water recharge during the vegetation restoration process in semi-arid regions of northern China. Hydrology Earth System Sci. 24, 5875–5890. doi: 10.5194/hess-24-5875-2020

Crossref Full Text | Google Scholar

Cui, Y., Pan, C., Ma, L., and Sun, Z. (2023). Comparing water uptake patterns of two tree species using stable isotopes on the loess plateau, China. Hydrological Processes 37, e15006. doi: 10.1002/hyp.15006

Crossref Full Text | Google Scholar

Deng, L., Yan, W., Zhang, Y., and Shangguan, Z. (2016). Severe depletion of soil moisture following land-use changes for ecological restoration: Evidence from northern China. For. Ecol. Manage. 366, 1–10. doi: 10.1016/j.foreco.2016.01.026

Crossref Full Text | Google Scholar

de Queiroz, M. G., da Silva, T. G. F., Zolnier, S., Jardim, A. M., da, R. F., de Souza, C. A. A., et al. (2020). Spatial and temporal dynamics of soil moisture for surfaces with a change in land use in the semi-arid region of Brazil. Catena 188, 104457. doi: 10.1016/j.catena.2020.104457

Crossref Full Text | Google Scholar

Dong, X., Lee, E., Gwak, Y., and Kim, S. (2018). Seasonal control of spatial distribution of soil moisture on a steep hillslope. Water Sci.Technol: A J. Int. Assoc. Water pollut. Res. 79 3, 556–565. doi: 10.2166/wcc.2018.173

PubMed Abstract | Crossref Full Text | Google Scholar

Dymond, S. F., Wagenbrenner, J. W., Keppeler, E. T., and Bladon, K. D. (2021). Dynamic hillslope soil moisture in a Mediterranean montane watershed. Water Resour. Res. 57, e2020WR029170. doi: 10.1029/2020WR029170

Crossref Full Text | Google Scholar

Fathololoumi, S., Vaezi, A. R., Firozjaei, M. K., and Biswas, A. (2021). Quantifying the effect of surface heterogeneity on soil moisture across regions and surface characteristic. J. Hydrology 596, 126132. doi: 10.1016/j.jhydrol.2021.126132

Crossref Full Text | Google Scholar

Gao, X., Wu, P., Zhao, X., Wang, J., and Shi, Y. (2014). Effects of land use on soil moisture variations in a semi-arid catchment: Implications for land and agricultural water management. Land Degradation Dev. 25, 163–172. doi: 10.1002/ldr.1156

Crossref Full Text | Google Scholar

Gao, Y. and Yang, P. (2023). Temporal and spatial distribution of soil water repellency in grassland soils and its relation to soil moisture, hydrophobic matter, and particle size. Sci. Total Environ. 904, 166700. doi: 10.1016/j.scitotenv.2023.166700

PubMed Abstract | Crossref Full Text | Google Scholar

Helman, D. and Mussery, A. (2020). Using landsat satellites to assess the impact of check dams built across erosive gullies on vegetation rehabilitation. Sci. Total Environ. 730, 138873. doi: 10.1016/j.scitotenv.2020.138873

PubMed Abstract | Crossref Full Text | Google Scholar

Hou, X., Jia, Y., Han, H., and Shao, M. (2018). Response of soil moisture to single-rainfall events under three vegetation types in the gully region of the loess plateau. Sustainability. 10, 3793. doi: 10.3390/SU10103793

Crossref Full Text | Google Scholar

Huang, X., Shi, Z., Zhu, H., Zhang, H., Ai, L., and Yin, W. (2016). Soil moisture dynamics within soil profiles and associated environmental controls. Catena 136, 189–196. doi: 10.1016/J.CATENA.2015.01.014

Crossref Full Text | Google Scholar

Jia, Y.-H. and Shao, M.-A. (2014). Dynamics of deep soil moisture in response to vegetational restoration on the loess plateau of China. J Hydrol. 519, 523–531. doi: 10.1016/j.jhydrol.2014.07.043

Crossref Full Text | Google Scholar

Jia, X., Shao, M., Zhu, Y., and Luo, Y. (2017). Soil moisture decline due to afforestation across the loess plateau, China. J. Hydrology 546, 113–122. doi: 10.1016/j.jhydrol.2017.01.011

Crossref Full Text | Google Scholar

Jian, S., Zhao, C., Fang, S., and Yu, K. (2015). Effects of different vegetation restoration on soil water storage and water balance in the Chinese loess plateau. Agric. For. Meteorology 206, 85–96. doi: 10.1016/j.agrformet.2015.03.009

Crossref Full Text | Google Scholar

Jin, Z., Guo, L., Lin, H., Wang, Y., Yu, Y., Chu, G., et al. (2018). Soil moisture response to rainfall on the Chinese loess plateau after a long-term vegetation rehabilitation. Hydrological Processes 32, 1738–1754. doi: 10.1002/hyp.13143

Crossref Full Text | Google Scholar

Li, B., Li, P., Zhang, W., Ji, J., Liu, G., and Xu, M. (2021). Deep soil moisture limits the sustainable vegetation restoration in arid and semi-arid Loess Plateau. Geoderma 399, 115122. doi: 10.1016/j.geoderma.2021.115122

Crossref Full Text | Google Scholar

Liu, X., Cai, L., Li, M., Yan, Y., Chen, H., and Wang, F. (2024a). Why does afforestation policy lead to a drying trend in soil moisture on the loess plateau? Sci. Total Environ. 953, 175912. doi: 10.1016/j.scitotenv.2024.175912

PubMed Abstract | Crossref Full Text | Google Scholar

Liu, X., Jiao, L., Cheng, D., Liu, J., Li, Z., Li, Z., et al. (2024b). Light thinning effectively improves forest soil water replenishment in water-limited areas: Observational evidence from robinia pseudoacacia plantations on the loess plateau, China. J. Hydrology 637, 131408. doi: 10.1016/j.jhydrol.2024.131408

Crossref Full Text | Google Scholar

Liu, Y., Li, S., Chen, F., Yang, S., and Chen, X. (2010). Soil water dynamics and water use efficiency in spring maize (zea mays L.) fields subjected to different water management practices on the loess plateau, China. Agric. Water Manage. 97, 769–775. doi: 10.1016/j.agwat.2010.01.010

Crossref Full Text | Google Scholar

Liu, Y., Miao, H., Huang, Z., Cui, Z., He, H., Zheng, J., et al. (2018). Soil water depletion patterns of artificial forest species and ages on the loess plateau (China). For. Ecol. Manage. 417, 137–143. doi: 10.1016/j.foreco.2018.03.005

Crossref Full Text | Google Scholar

Luan, J., Miao, P., Tian, X., Li, X., Ma, N., Xu, Z., et al. (2022). Separating the impact of check dams on runoff from climate and vegetation changes. J. Hydrology 614, 128565. doi: 10.1016/j.jhydrol.2022.128565

Crossref Full Text | Google Scholar

Mei, X. and Ma, L. (2022). Effect of afforestation on soil water dynamics and water uptake under different rainfall types on the loess hillslope. Catena 213, 106216. doi: 10.1016/j.catena.2022.106216

Crossref Full Text | Google Scholar

Montenegro, S. and Ragab, R. (2012). Impact of possible climate and land use changes in the semi arid regions: A case study from North Eastern Brazil. J. Hydrology 434–435, 55–68. doi: 10.1016/j.jhydrol.2012.02.036

Crossref Full Text | Google Scholar

O’Donnell, F. C., Donager, J., Sankey, T., Masek Lopez, S., and Springer, A. E. (2021). Vegetation structure controls on snow and soil moisture in restored ponderosa pine forests. Hydrological Processes 35, e14432. doi: 10.1002/hyp.14432

Crossref Full Text | Google Scholar

Pal, M. and Maity, R. (2021). Assimilation of remote sensing based surface soil moisture to develop a spatially varying vertical soil moisture profile database for entire Indian mainland. J. Hydrology 601, 126807. doi: 10.1016/j.jhydrol.2021.126807

Crossref Full Text | Google Scholar

Sharma, A., Patel, P. L., and Sharma, P. J. (2022). Influence of climate and land-use changes on the sensitivity of SWAT model parameters and water availability in a semi-arid river basin. Catena 215, 106298. doi: 10.1016/j.catena.2022.106298

Crossref Full Text | Google Scholar

Shen, Q., Gao, G., Fu, B., and Lü, Y. (2014). Soil water content variations and hydrological relations of the cropland-treebelt-desert land use pattern in an oasis-desert ecotone of the heihe river basin, China. Catena 123, 52–61. doi: 10.1016/j.catena.2014.07.002

Crossref Full Text | Google Scholar

Shi, Z., Zhu, H., Chen, J., Fang, N., and Ai, L. (2012). Spatial heterogeneity of soil moisture and its relationships with environmental factors at small catchment level. J. Appl. Ecol. 23 4, 889–895. doi: 10.13287/j.1001-9332.2012.0124

PubMed Abstract | Crossref Full Text | Google Scholar

Tijdeman, E. and Menzel, L. (2021). The development and persistence of soil moisture stress during drought across southwestern Germany. Hydrology Earth System Sci. 25, 2009–2025. doi: 10.5194/hess-25-2009-2021

Crossref Full Text | Google Scholar

Tuure, J., Räsänen, M., Hautala, M., Pellikka, P., Mäkelä, P. S. A., and Alakukku, L. (2021). Plant residue mulch increases measured and modelled soil moisture content in the effective root zone of maize in semi-arid Kenya. Soil Tillage Res. 209, 104945. doi: 10.1016/j.still.2021.104945

Crossref Full Text | Google Scholar

Wang, L., Dalabay, N., Lu, P., and Wu, F. (2017). Effects of tillage practices and slope on runoff and erosion of soil from the loess plateau, China, subjected to simulated rainfall. Soil Tillage Res. 166, 147–156. doi: 10.1016/j.still.2016.09.007

Crossref Full Text | Google Scholar

Wang, Y., Gao, L., Li, J., and Peng, X. (2024a). Responses of deep soil moisture to direct rainfall and groundwater in the red soil critical zone: A four-stage pattern. J. Hydrology 632, 130864. doi: 10.1016/j.jhydrol.2024.130864

Crossref Full Text | Google Scholar

Wang, Y., Hu, W., Sun, H., Zhao, Y., Zhang, P., Li, Z., et al. (2024b). Soil moisture decline in China’s monsoon loess critical zone: More a result of land-use conversion than climate change. Proc. Natl. Acad. Sci. 121, e2322127121. doi: 10.1073/pnas.2322127121

PubMed Abstract | Crossref Full Text | Google Scholar

Wang, Y., Jiang, Y., Zhao, Y., Li, X., Xie, K., Yan, T., et al. (2024). Modeling the impact of long-term land use changes on deep soil hydrological processes in the loess plateau, China. J. Hydrology 643, 131944. doi: 10.1016/j.jhydrol.2024.131944

Crossref Full Text | Google Scholar

Wang, J., Liu, Z., Gao, J., Emanuele, L., Ren, Y., Shao, M., et al. (2021). The grain for green project eliminated the effect of soil erosion on organic carbon on China’s loess plateau between 1980 and 2008. Agric. Ecosyst. Environ. 322, 107636. doi: 10.1016/j.agee.2021.107636

Crossref Full Text | Google Scholar

Wang, X.-Z., Jiao, F., Liu, Y.-X., Zhu, L.-T., and Lin, K. (2012). Relationships between soil moisture and environmental factors at different spatial scales. Chinese J Ecol 312, 319–323. doi: 10.13292/j.1000-4890.2012.0038

Crossref Full Text | Google Scholar

Wang, Y., Sun, H., and Zhao, Y. (2019). Characterizing spatial-temporal patterns and abrupt changes in deep soil moisture across an intensively managed watershed. Geoderma 341, 181–194. doi: 10.1016/j.geoderma.2019.01.044

Crossref Full Text | Google Scholar

Wang, S., Yang, M., Gao, X., Li, B., Cai, Y., Li, C., et al. (2024). Ecohydrological response to deep soil desiccation in a semiarid apple orchard. Agric. For. Meteorology 354, 110089. doi: 10.1016/j.agrformet.2024.110089

Crossref Full Text | Google Scholar

Wu, W., Li, H., Feng, H., Si, B., Chen, G., Meng, T., et al. (2021). Precipitation dominates the transpiration of both the economic forest (malus pumila) and ecological forest (robinia pseudoacacia) on the loess plateau after about 15 years of water depletion in deep soil. Agric. For. Meteorology 297, 108244. doi: 10.1016/j.agrformet.2020.108244

Crossref Full Text | Google Scholar

Wyatt, B. M., Ochsner, T. E., and Zou, C. B. (2021). Estimating root zone soil moisture across diverse land cover types by integrating in-situ and remotely sensed data. Agric. For. Meteorology 307, 108471. doi: 10.1016/j.agrformet.2021.108471

Crossref Full Text | Google Scholar

Xiao, L., Xue, S., Liu, G., and Zhang, C. (2014). Soil moisture variability under different land uses in the zhifanggou catchment of the loess plateau, China. Arid Land Res. Manage. 28, 274–290. doi: 10.1080/15324982.2013.860500

Crossref Full Text | Google Scholar

Xu, G., Huang, M., Li, P., Li, Z., and Wang, Y. (2021). Effects of land use on spatial and temporal distribution of soil moisture within profiles. Environ. Earth Sci. 80, 128. doi: 10.1007/s12665-021-09464-2

Crossref Full Text | Google Scholar

Yang, L., Chen, L., Wei, W., Yu, Y., and Zhang, H. (2014). Comparison of deep soil moisture in two re-vegetation watersheds in semi-arid regions. J. Hydrology 513, 314–321. doi: 10.1016/j.jhydrol.2014.03.049

Crossref Full Text | Google Scholar

Yang, L., Wei, W., Chen, L., and Mo, B. (2012). Response of deep soil moisture to land use and afforestation in the semi-arid loess plateau, China. J. Hydrology 475, 111–122. doi: 10.1016/j.jhydrol.2012.09.041

Crossref Full Text | Google Scholar

Yao, Y., Wang, X., Zeng, Z., Liu, Y., Peng, S., Zhu, Z., et al. (2016). The effect of afforestation on soil moisture content in northeastern China. PloS One 11, e0160776. doi: 10.1371/journal.pone.0160776

PubMed Abstract | Crossref Full Text | Google Scholar

Ye, L., Fang, L., Shi, Z., Deng, L., and Tan, W. (2019). Spatio-temporal dynamics of soil moisture driven by ‘grain for green’ program on the loess plateau, China. Agric. Ecosyst. Environ. 269, 204–214. doi: 10.1016/j.agee.2018.10.006

Crossref Full Text | Google Scholar

Yu, Y., Hua, T., Chen, L., Zhang, Z., and Pereira, P. (2024). Divergent changes in vegetation greenness, productivity, and rainfall use efficiency are characteristic of ecological restoration towards high-quality development in the Yellow River basin, China. Engineering 34, 109–119. doi: 10.1016/j.eng.2023.07.012

Crossref Full Text | Google Scholar

Yu, B., Liu, G., Liu, Q., Huang, C., and Li, H. (2019). Effects of topographic domain and land use on spatial variability of deep soil moisture in the semi-arid loess plateau of China. Hydrology Res. 50, 1281–1292. doi: 10.2166/nh.2019.221

Crossref Full Text | Google Scholar

Yu, B., Liu, G., Liu, Q., Wang, X., Feng, J., and Huang, C. (2018). Soil moisture variations at different topographic domains and land use types in the semi-arid loess plateau, China. Catena 165, 125–132. doi: 10.1016/j.catena.2018.01.020

Crossref Full Text | Google Scholar

Yu, Y., Zhao, W., Martinez-Murillo, J. F., and Pereira, P. (2020). Loess plateau: From degradation to restoration. Sci. Total Environ. 738, 140206. doi: 10.1016/j.scitotenv.2020.140206

PubMed Abstract | Crossref Full Text | Google Scholar

Zhang, Y., Feng, S., Wang, J., Chen, M., Wang, K., Liu, C., et al. (2024). Assessing the change in soil water deficit characteristics from grassland to forestland on the loess plateau. Ecol. Indic. 167, 112616. doi: 10.1016/j.ecolind.2024.112616

Crossref Full Text | Google Scholar

Zhang, Q., Jia, X., Zhao, C., and Shao, M. (2018). Revegetation with artificial plants improves topsoil hydrological properties but intensifies deep-soil drying in northern loess plateau, China. J. Arid Land 10, 335–346. doi: 10.1007/s40333-018-0007-0

Crossref Full Text | Google Scholar

Zhou, Z., Wang, Y., An, Z., Li, R., Xu, Y., Zhang, P., et al. (2022). Deep root information “hidden in the dark”: A case study on the 21-m soil profile of robinia pseudoacacia in the critical zone of the chinese loess plateau. Catena 213, 106121. doi: 10.1016/j.catena.2022.106121

Crossref Full Text | Google Scholar

Zhou, J., Wang, Y., Li, R., He, H., Sun, H., Zhou, Z., et al. (2024). Response of deep soil water deficit to afforestation, soil depth, and precipitation gradient. Agric. For. Meteorology 352, 110024. doi: 10.1016/j.agrformet.2024.110024

Crossref Full Text | Google Scholar

Keywords: soil moisture, vegetation restoration, vertical distribution, spatial variability, Loess Plateau

Citation: Zhu H, Wang Z, Zhao J, Lin J, Qian S, Wang L, Yu Y and Cavalli M (2025) The hilly-gully watershed exhibits distinct deep soil moisture characteristics: a comparative study of paired watersheds in the Chinese Loess Plateau. Front. Plant Sci. 16:1668310. doi: 10.3389/fpls.2025.1668310

Received: 17 July 2025; Accepted: 05 September 2025;
Published: 19 September 2025.

Edited by:

Zhi-Qiang Zhang, Yunnan University, China

Reviewed by:

Shi-jian Yang, Yunnan University, China
Ashley Webb, WaterNSW, Australia

Copyright © 2025 Zhu, Wang, Zhao, Lin, Qian, Wang, Yu and Cavalli. This is an open-access article distributed under the terms of the Creative Commons Attribution License (CC BY). The use, distribution or reproduction in other forums is permitted, provided the original author(s) and the copyright owner(s) are credited and that the original publication in this journal is cited, in accordance with accepted academic practice. No use, distribution or reproduction is permitted which does not comply with these terms.

*Correspondence: Yang Yu, dGhlb2RvcmV5eUBnbWFpbC5jb20=

Disclaimer: All claims expressed in this article are solely those of the authors and do not necessarily represent those of their affiliated organizations, or those of the publisher, the editors and the reviewers. Any product that may be evaluated in this article or claim that may be made by its manufacturer is not guaranteed or endorsed by the publisher.